INO80 complex regulates the G1-to-S transcriptional wave through MBF

In eukaryotic cells a key regulatory step of cell cycle occurs in late G1, which has been termed “Start” in yeast cells. At this point, cells determine whether they will go to through a new round of proliferation, or choose alternative pathways; cell cycle arrest or sexual differentiation. In fissio...

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Detalles Bibliográficos
Autor: Knezevic, Iva
Tipo de recurso: tesis doctoral
Estado:Versión publicada
Fecha de publicación:2015
País:España
Institución:CBUC, CESCA
Repositorio:TDR. Tesis Doctorales en Red
OAI Identifier:oai:www.tdx.cat:10803/586378
Acceso en línea:http://hdl.handle.net/10803/586378
Access Level:acceso abierto
Palabra clave:S. pombe
MBF
INO80
Transcription factor
DNA damage
Les4
Histona H2A.Z
Nucleosomas
Transcripción
Ciclo celular
576
Descripción
Sumario:In eukaryotic cells a key regulatory step of cell cycle occurs in late G1, which has been termed “Start” in yeast cells. At this point, cells determine whether they will go to through a new round of proliferation, or choose alternative pathways; cell cycle arrest or sexual differentiation. In fission yeast, the MBF complex controls the transcriptional activation of some genes of G1/S transcriptional wave, including cdc18, cdt1 (necessary to prevent onset of mitosis before of completion of DNA synthesis), and cdc22 (ribonucleotide reductase). Control of MBF activity is essential for normal cell cycle progression. It has been found that MBF complex is bound to the promoter of its target genes throughout the cell cycle, implicating that MBF activity is not regulated by pure binding to DNA. We purified novel inetractor of MBF complex, INO80 complex. Here we demonstrate that INO80 regulates cell cycle genes through MBF, and that proper acetylation of histone variant H2A.Z is crucial for MBF dependent transcription.