Histone H1 regulates non-coding RNA turnover on chromatin in a m6A-dependent manner

Linker histones are highly abundant chromatin-associated proteins with well-established structural roles in chromatin and as general transcriptional repressors. In addition, it has been long proposed that histone H1 exerts context-specific effects on gene expression. Here, we identify a function of...

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Detalles Bibliográficos
Autores: Fernández-Justel, José Miguel, Santa-María, Cristina, Martín-Vírgala, Sara, Ramesh, Shreya, Ferrera-Lagoa, Alberto, Salinas Pena, Mónica, Isoler-Alcaráz, Javier, Maslon, Magdalena M., Jordan, Albert, Cáceres, Javier F., Gómez, María
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2022
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/281158
Acceso en línea:http://hdl.handle.net/10261/281158
Access Level:acceso abierto
Palabra clave:Histone H1
lncRNAs
Chromatin RNAs
Replication-transcription conflicts
m6A
Replicative stress
R-loops
Chromatin conformation
mESCs
Descripción
Sumario:Linker histones are highly abundant chromatin-associated proteins with well-established structural roles in chromatin and as general transcriptional repressors. In addition, it has been long proposed that histone H1 exerts context-specific effects on gene expression. Here, we identify a function of histone H1 in chromatin structure and transcription using a range of genomic approaches. In the absence of histone H1, there is an increase in the transcription of non-coding RNAs, together with reduced levels of m6A modification leading to their accumulation on chromatin and causing replication-transcription conflicts. This strongly suggests that histone H1 prevents non-coding RNA transcription and regulates non-coding transcript turnover on chromatin. Accordingly, altering the m6A RNA methylation pathway rescues the replicative phenotype of H1 loss. This work unveils unexpected regulatory roles of histone H1 on non-coding RNA turnover and m6A deposition, highlighting the intimate relationship between chromatin conformation, RNA metabolism, and DNA replication to maintain genome performance.