Integrin and CD3/TCR activation are regulated by the scaffold protein AKAP450

During antigen recognition by T cells, membrane receptors and cytoskeletal molecules form a specialized structure at the T cell–antigen-presenting cell junction called the immune synapse (IS). We report a role for the scaffolding protein A-kinase anchoring protein-450 (AKAP450), a member of the A-ki...

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Detalhes bibliográficos
Autores: Robles-Valero, Javier, Martín-Cófreces, Noa, Lamana Domínguez, Amalia, Macdonald, Stephen, Volkov, Yuri, Sánchez-Madrid, Francisco
Formato: artículo
Fecha de publicación:2010
País:España
Recursos:Universidad Complutense de Madrid (UCM)
Repositorio:Docta Complutense
Idioma:inglés
OAI Identifier:oai:docta.ucm.es:20.500.14352/94342
Acesso em linha:https://hdl.handle.net/20.500.14352/94342
Access Level:acceso abierto
Palavra-chave:576.3
577.112
577.27
Integrin
CD3/TCRactivation
Protein AKAP450
Biología celular (Biología)
Bioquímica (Biología)
Inmunología
2407 Biología Celular
2403 Bioquímica
2412 Inmunología
Descrição
Resumo:During antigen recognition by T cells, membrane receptors and cytoskeletal molecules form a specialized structure at the T cell–antigen-presenting cell junction called the immune synapse (IS). We report a role for the scaffolding protein A-kinase anchoring protein-450 (AKAP450), a member of the A-kinase anchoring protein family, in IS formation and T-cell signaling in antigen- and superantigen-dependent T-cell activation. Suppression of AKAP450 by overexpression of a dominant-negative form or siRNA knockdown disrupted the positioning and conformational activation of lymphocyte function-associated antigen 1 at the IS and impaired associated signaling events, including phosphorylation of phospholipase C-γ1 and protein kinase C-θ. AKAP450 was also required for correct activation and phosphorylation of CD3, LAT, and Vav1, key T-cell receptor-activated intracellular signaling molecules. Consistently, antigen-triggered reorientation of the microtubule-organizing center at the IS and interleukin-2 secretion were diminished in AKAP450-disrupted T cells. These results indicate key roles for AKAP450 in the organization and activation of receptor molecules at the IS during T-cell signaling events.