Max1 links MBF dependent transcription upon completion of DNA synthesis in fission yeast

When DNA replication is challenged, cells activate a DNA synthesis checkpoint blocking cell cycle progression until they are able to overcome the replication defects. In fission yeast, Cds1 is the effector kinase of this checkpoint, inhibiting M phase entry, stabilizing stalled replication forks and...

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Detalles Bibliográficos
Autor: Gómez Escoda, Blanca
Tipo de recurso: tesis doctoral
Estado:Versión publicada
Fecha de publicación:2010
País:España
Institución:CBUC, CESCA
Repositorio:TDR. Tesis Doctorales en Red
OAI Identifier:oai:www.tdx.cat:10803/7223
Acceso en línea:http://www.tdx.cat/TDX-0207111-150509
http://hdl.handle.net/10803/7223
Access Level:acceso abierto
Palabra clave:stress
kinase
phosphorylation
DNA damage
Checkpoint
Cell Cycle
transcription factor
transcription
replication
yeast
estrés
kinasa
fosforilación
daño en DNA
checkpoint
ciclo celular
factor de transcripción
transcripción
replicación
Schizosaccharomyces pombe
levadura
577
Descripción
Sumario:When DNA replication is challenged, cells activate a DNA synthesis checkpoint blocking cell cycle progression until they are able to overcome the replication defects. In fission yeast, Cds1 is the effector kinase of this checkpoint, inhibiting M phase entry, stabilizing stalled replication forks and triggering transcriptional activation of S-phase genes; the molecular basis of this last effect remains largely unknown. The MBF complex controls the transcription of S-phase genes. We have purified novel interactors of the MBF complex and among them we have identified the repressor Max1. When the DNA synthesis checkpoint is activated, Max1 is phosphorylated by Cds1 resulting in the abrogation of its binding to MBF. As a consequence, MBF-dependent transcription is maintained active until cells are able to overcome this challenge.