Autophosphorylation and transphosphorylation mechanisms in bacterial two component systems

Two-component signaling circuits (TCS) allow bacteria to detect environmental cues and to produce adaptive responses. These signaling systems are based on autophosphorylation and phosphoryl-group transfers between histidine and aspartate containing sensor kinase and response regulator proteins. Upon...

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Bibliographic Details
Authors: Terán-Melo, Juan Luis, Rodríguez-Rangel, Claudia, Georgellis, Dimitris, Álvarez, Adrián F.
Format: article
Status:Published version
Publication Date:2019
Country:México
Institution:UNIVERSIDAD NACIONAL AUTÓNOMA DE MÉXICO
Repository:TIP Revista especializada en ciencias químico-biológicas
Language:Spanish
OAI Identifier:oai:ojs.ojs.escire.net:article/162
Online Access:http://tip.zaragoza.unam.mx/index.php/tip/article/view/162
Access Level:Open access
Keyword:sensor kinase; autophosphorylation; transphosphorylation; intermolecular; intramolecular
cinasa sensora; autofosforilación; transfosforilación; intermolecular; intramolecular
Description
Summary:Two-component signaling circuits (TCS) allow bacteria to detect environmental cues and to produce adaptive responses. These signaling systems are based on autophosphorylation and phosphoryl-group transfers between histidine and aspartate containing sensor kinase and response regulator proteins. Upon reception of a specific stimulus the sensor kinase protein autophosphorylates, by either an inter- or intra-molecular reaction, and transphosphorylates its cognate response regulator, which, typically, acts as a transcriptional regulator, thereby triggering physiological responses. Frequently, in the absence of the stimulus, the sensor kinase proteins are responsible for the dephosphorylation of their cognate response regulators. Furthermore, a group of sensor kinases have additional functional domains that are involved in a forward phosphorelay for signal transmission and in a reverse phosphorelay for signal decay. As is the case of the autophosphorylation reaction, the phosphoryl-group transfers involved in the forward and reverse phosphorelay can occur either intra- or inter-molecularly. In this review, we highlight some important features of bacterial TCS, with special emphasis on the autophosphorylation and phosphoryl-group transfer events.