Atp: The crucial component of secretory vesicles

The colligative properties of ATP and catecholamines demonstrated in vitro are thought to be responsible for the extraordinary accumulation of solutes inside chromaffin cell secretory vesicles, although this has yet to be demonstrated in living cells. Because functional cells cannot be deprived of A...

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Detalles Bibliográficos
Autores: Estévez Herrera, Judith, Domínguez, Natalia, Pardo, Marta R., González Santana, Ayoze, Westhead, Edward W., Borges Jurado, Ricardo, Machado Ponce, José David
Tipo de recurso: artículo
Fecha de publicación:2016
País:España
Institución:Universidad de La Laguna (ULL)
Repositorio:RIULL. Repositorio Institucional de la Universidad de La Laguna
OAI Identifier:oai:riull.ull.es:915/41334
Acceso en línea:http://riull.ull.es/xmlui/handle/915/41334
Access Level:acceso abierto
Palabra clave:exocytosis
purines
quantum size
secretory vesicles
VNUT
Descripción
Sumario:The colligative properties of ATP and catecholamines demonstrated in vitro are thought to be responsible for the extraordinary accumulation of solutes inside chromaffin cell secretory vesicles, although this has yet to be demonstrated in living cells. Because functional cells cannot be deprived of ATP, we have knocked down theexpression ofthe vesicular nucleotide carrier, the VNUT,to show that a reduction in vesicular ATP is accompanied by a drastic fall in the quantal release of catecholamines. This phenomenon is particularly evident in newly synthesized vesicles, which we show are the first to be released. Surprisingly, we find that inhibiting VNUT expression also reduces the frequency of exocytosis, whereas the overexpression of VNUT drastically increases the quantal size of exocytotic events. To our knowledge, our data provide the first demonstration that ATP, in addition to serving as an energy source and purinergic transmitter, is an essential element in the concentration of catecholamines in secretory vesicles. In this way, cells can use ATP to accumulate neurotransmitters and other secreted substances at high concentrations, supporting quantal transmission.