Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification

[EN] 1. Intrinsic water use efficiency (WUEi) reflects the trade-off between photo- synthetic carbon gain and water loss through stomatal conductance and is key for understanding dryland plant responses to climate change. Stipa tenacissima is a perennial tussock C3 grass with an opportunistic, droug...

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Autores: Ren, Wei, García Palacios, Pablo, Soliveres Codina, Santiago, Prieto Aguilar, Iván, Maestre Gil, Fernando Tomás, Querejeta Mercader, José Ignacio
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2024
País:España
Institución:Universidad de León
Repositorio:BULERIA. Repositorio Institucional de la Universidad de León
OAI Identifier:oai:buleria.unileon.es:10612/20866
Acceso en línea:https://besjournals.onlinelibrary.wiley.com/doi/epdf/10.1111/1365-2435.14518
https://hdl.handle.net/10612/20866
Access Level:acceso abierto
Palabra clave:Ecología. Medio ambiente
Climate aridification
Drought avoidance
Drylands
Foliar nutrients
Intrinsic water use efficiency
Stipa tenacissima
Stomatal regulation
Water and nitrogen co-limitation
2508.13 Humedad del Suelo
2511.02 Biología de Suelos
2417.13 Ecología Vegetal
2502.03 Bioclimatología
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oai_identifier_str oai:buleria.unileon.es:10612/20866
network_acronym_str ES
network_name_str España
repository_id_str
dc.title.none.fl_str_mv Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
title Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
spellingShingle Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
Ren, Wei
Ecología. Medio ambiente
Climate aridification
Drought avoidance
Drylands
Foliar nutrients
Intrinsic water use efficiency
Stipa tenacissima
Stomatal regulation
Water and nitrogen co-limitation
2508.13 Humedad del Suelo
2511.02 Biología de Suelos
2417.13 Ecología Vegetal
2502.03 Bioclimatología
title_short Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
title_full Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
title_fullStr Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
title_full_unstemmed Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
title_sort Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridification
dc.creator.none.fl_str_mv Ren, Wei
García Palacios, Pablo
Soliveres Codina, Santiago
Prieto Aguilar, Iván
Maestre Gil, Fernando Tomás
Querejeta Mercader, José Ignacio
author Ren, Wei
author_facet Ren, Wei
García Palacios, Pablo
Soliveres Codina, Santiago
Prieto Aguilar, Iván
Maestre Gil, Fernando Tomás
Querejeta Mercader, José Ignacio
author_role author
author2 García Palacios, Pablo
Soliveres Codina, Santiago
Prieto Aguilar, Iván
Maestre Gil, Fernando Tomás
Querejeta Mercader, José Ignacio
author2_role author
author
author
author
author
dc.contributor.none.fl_str_mv Ecologia
Facultad de Ciencias Biologicas y Ambientales
dc.subject.none.fl_str_mv Ecología. Medio ambiente
Climate aridification
Drought avoidance
Drylands
Foliar nutrients
Intrinsic water use efficiency
Stipa tenacissima
Stomatal regulation
Water and nitrogen co-limitation
2508.13 Humedad del Suelo
2511.02 Biología de Suelos
2417.13 Ecología Vegetal
2502.03 Bioclimatología
topic Ecología. Medio ambiente
Climate aridification
Drought avoidance
Drylands
Foliar nutrients
Intrinsic water use efficiency
Stipa tenacissima
Stomatal regulation
Water and nitrogen co-limitation
2508.13 Humedad del Suelo
2511.02 Biología de Suelos
2417.13 Ecología Vegetal
2502.03 Bioclimatología
description [EN] 1. Intrinsic water use efficiency (WUEi) reflects the trade-off between photo- synthetic carbon gain and water loss through stomatal conductance and is key for understanding dryland plant responses to climate change. Stipa tenacissima is a perennial tussock C3 grass with an opportunistic, drought- avoiding water use strategy that dominates arid and semiarid steppes across the western Mediterranean region. However, its ecophysiological responses to aridification and woody shrub encroachment, a major land-use change in drylands worldwide, are not well-understood. 2. We investigated the variations in leaf stable isotopes (δ18 O, δ13 C, δ15 N), nutrient concentrations (N, P, K), and culm water content and isotopic composition (δ18 O,δ2H) of paired pure-grass and shrub-encroached S. tenacissima steppes along a 350 km aridity gradient in Spain (10 sites, 160 individuals). 3. Culm water isotopes revealed that S. tenacissima is a shallow-rooted grass that depends heavily on recent rainwater for water uptake, which may render it vulnerable to increasingly irregular rainfall combined with faster topsoil drying under climate warming and aridification. With increasing aridity, S. tenacissima enhanced leaf-level WUEi through more stringent stomatal regulation of plant water flux and carbon assimilation (higher δ13C and δ18 O), reaching exceptionally high δ13C values (−23‰ to −21‰) at the most arid steppes. Foliar N concentration was remarkably low across sites regardless of woody shrub encroachment, evidencing severe water and N co-limitation of photosynthesis and productivity. Shrub encroachment decreased leaf P and K but did not affect S. tenacissima water status. Perennial grass cover decreased markedly with both declining winter rainfall and shrub encroachment suggesting population-level rather than individual-level responses of S. tenacissima to these changes.4. The fundamental physiological constraints of photosynthetic C3 metabolism com-bined with low foliar N content may hamper the ability of S. tenacissima and other drought-avoider species with shallow roots to achieve further adaptive improve-ments in WUEi under increasing climatic stress. A drought-avoiding water use strategy based on early stomatal closure and photosynthesis suppression during prolonged rainless periods may thus compromise the capacity of semiarid S. tena-cissima steppes to maintain perennial grass cover, sustain productivity and cope with ongoing climate aridification at the drier parts of their current distribution
publishDate 2024
dc.date.none.fl_str_mv 2024
dc.type.none.fl_str_mv info:eu-repo/semantics/article
info:eu-repo/semantics/publishedVersion
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv https://besjournals.onlinelibrary.wiley.com/doi/epdf/10.1111/1365-2435.14518
https://hdl.handle.net/10612/20866
url https://besjournals.onlinelibrary.wiley.com/doi/epdf/10.1111/1365-2435.14518
https://hdl.handle.net/10612/20866
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
dc.relation.none.fl_str_mv info:eu-repo/grantAgreement/MICINN//EUR2022-134048
info:eu-repo/grantAgreement/MINECO/Programa Estatal de Fomento de la Investigación Científica y Técnica de Excelencia/CGL2013-48753-R
info:eu-repo/grantAgreement/MICINN/Programa Nacional de Investigación Fundamental/CGL2010–21064
info:eu-repo/grantAgreement/MEC//AGL2006-11234
info:eu-repo/grantAgreement/AEI/Programa Estatal de I+D+i Orientada a los Retos de la Sociedad/PID2020-113021RA-I00
info:eu-repo/grantAgreemen/AEI/Programa Estatal de Generación de Conocimiento y Fortalecimiento Científico y Tecnológico del Sistema de I+D+i/PID2019-107382RB-I00/ES/Diversidad funcional de estrategias de uso de agua y nutrientes por las plantas en ecosistemas mediterráneos y secos
info:eu-repo/grantAgreement/AEI/Programa Estatal de I+D+i Orientada a los Retos de la Sociedad/RTI2018- 098895-A-100
dc.rights.none.fl_str_mv http://creativecommons.org/licenses/by-nc-nd/4.0/
info:eu-repo/semantics/openAccess
rights_invalid_str_mv http://creativecommons.org/licenses/by-nc-nd/4.0/
eu_rights_str_mv openAccess
dc.publisher.none.fl_str_mv British Ecological Society
John Wiley & Sons Ltd
publisher.none.fl_str_mv British Ecological Society
John Wiley & Sons Ltd
dc.source.none.fl_str_mv reponame:BULERIA. Repositorio Institucional de la Universidad de León
instname:Universidad de León
instname_str Universidad de León
reponame_str BULERIA. Repositorio Institucional de la Universidad de León
collection BULERIA. Repositorio Institucional de la Universidad de León
repository.name.fl_str_mv
repository.mail.fl_str_mv
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spelling Pushing the limits of C3 intrinsic water use efficiency in Mediterranean semiarid steppes: responses of a drought‐avoider perennial grass to climate aridificationRen, WeiGarcía Palacios, PabloSoliveres Codina, SantiagoPrieto Aguilar, IvánMaestre Gil, Fernando TomásQuerejeta Mercader, José IgnacioEcología. Medio ambienteClimate aridificationDrought avoidanceDrylandsFoliar nutrientsIntrinsic water use efficiencyStipa tenacissimaStomatal regulationWater and nitrogen co-limitation2508.13 Humedad del Suelo2511.02 Biología de Suelos2417.13 Ecología Vegetal2502.03 Bioclimatología[EN] 1. Intrinsic water use efficiency (WUEi) reflects the trade-off between photo- synthetic carbon gain and water loss through stomatal conductance and is key for understanding dryland plant responses to climate change. Stipa tenacissima is a perennial tussock C3 grass with an opportunistic, drought- avoiding water use strategy that dominates arid and semiarid steppes across the western Mediterranean region. However, its ecophysiological responses to aridification and woody shrub encroachment, a major land-use change in drylands worldwide, are not well-understood. 2. We investigated the variations in leaf stable isotopes (δ18 O, δ13 C, δ15 N), nutrient concentrations (N, P, K), and culm water content and isotopic composition (δ18 O,δ2H) of paired pure-grass and shrub-encroached S. tenacissima steppes along a 350 km aridity gradient in Spain (10 sites, 160 individuals). 3. Culm water isotopes revealed that S. tenacissima is a shallow-rooted grass that depends heavily on recent rainwater for water uptake, which may render it vulnerable to increasingly irregular rainfall combined with faster topsoil drying under climate warming and aridification. With increasing aridity, S. tenacissima enhanced leaf-level WUEi through more stringent stomatal regulation of plant water flux and carbon assimilation (higher δ13C and δ18 O), reaching exceptionally high δ13C values (−23‰ to −21‰) at the most arid steppes. Foliar N concentration was remarkably low across sites regardless of woody shrub encroachment, evidencing severe water and N co-limitation of photosynthesis and productivity. Shrub encroachment decreased leaf P and K but did not affect S. tenacissima water status. Perennial grass cover decreased markedly with both declining winter rainfall and shrub encroachment suggesting population-level rather than individual-level responses of S. tenacissima to these changes.4. The fundamental physiological constraints of photosynthetic C3 metabolism com-bined with low foliar N content may hamper the ability of S. tenacissima and other drought-avoider species with shallow roots to achieve further adaptive improve-ments in WUEi under increasing climatic stress. A drought-avoiding water use strategy based on early stomatal closure and photosynthesis suppression during prolonged rainless periods may thus compromise the capacity of semiarid S. tena-cissima steppes to maintain perennial grass cover, sustain productivity and cope with ongoing climate aridification at the drier parts of their current distributionSISpanish Ministry of Science and Innovation, Grant/Award Number: EUR2022-134048, PRX19/00301, PID2019-107382RB-I00, CGL2013- 48753-R, CGL2010-21064, AGL- 2006-11234, PID2020-113021RA-I00, RTI2018- 098895-A-100 and RYC- 2016-20604; Fundación BBVA, Grant/ Award Number: BIOCON06/105; Universidad Rey Juan Carlos, Grant/Award Number: URJC-RNT- 063-2; European Research Council, Grant/Award Number: 647038; Generalitat Valenciana, Grant/ Award Number: CIDEGENT/2018/041; National Natural Science Foundation of China, Grant/Award Number: 41801091; China Postdoctoral Science Foundation, Grant/Award Number: 2018M643542 and 2019T120868; Fundación Séneca, Grant/ Award Number: 20654/JLI/1British Ecological SocietyJohn Wiley & Sons LtdEcologiaFacultad de Ciencias Biologicas y Ambientales2024info:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionhttps://besjournals.onlinelibrary.wiley.com/doi/epdf/10.1111/1365-2435.14518https://hdl.handle.net/10612/20866reponame:BULERIA. Repositorio Institucional de la Universidad de Leóninstname:Universidad de LeónInglésinfo:eu-repo/grantAgreement/MICINN//EUR2022-134048info:eu-repo/grantAgreement/MINECO/Programa Estatal de Fomento de la Investigación Científica y Técnica de Excelencia/CGL2013-48753-Rinfo:eu-repo/grantAgreement/MICINN/Programa Nacional de Investigación Fundamental/CGL2010–21064info:eu-repo/grantAgreement/MEC//AGL2006-11234info:eu-repo/grantAgreement/AEI/Programa Estatal de I+D+i Orientada a los Retos de la Sociedad/PID2020-113021RA-I00info:eu-repo/grantAgreemen/AEI/Programa Estatal de Generación de Conocimiento y Fortalecimiento Científico y Tecnológico del Sistema de I+D+i/PID2019-107382RB-I00/ES/Diversidad funcional de estrategias de uso de agua y nutrientes por las plantas en ecosistemas mediterráneos y secosinfo:eu-repo/grantAgreement/AEI/Programa Estatal de I+D+i Orientada a los Retos de la Sociedad/RTI2018- 098895-A-100http://creativecommons.org/licenses/by-nc-nd/4.0/info:eu-repo/semantics/openAccessoai:buleria.unileon.es:10612/208662026-06-24T12:43:27Z
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