A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation

Actin cytoskeleton dynamics is essential for proper nervous system development and function. A conserved set of neuronal-specific microexons influences multiple aspects of neurobiology; however, their roles in regulating the actin cytoskeleton are unknown. Here, we study a microexon in DAAM1, a form...

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Autores: Poliński, Patryk, Miret-Cuesta, Marta, Zamora Moratalla, Alfonsa, Mantica, Federica, Cantero Recasens, Gerard, 1984-, Viana, Carlotta, Sabariego Navarro, Miguel, Normanno, Davide, Iñiguez, Luis P., Bonnal, Sophie, Gómez-Riera, Raúl, Fanlo-Ucar, Hugo, Yap, Dominic S., Martínez de Lagrán Cabredo, María, Fernández-Blanco, Álvaro, Rodríguez-Marin, Cristina, Permanyer, Jon, Fölsz, Orsolya, Domínguez-Sala, Eduardo, Sierra, Cesar, Wojnacki, José, Musoles Lleo, Juan Luis, Cosma, Maria Pia, Muñoz López, Francisco José, 1964-, Dierssen, Mara, Irimia Martínez, Manuel
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2025
País:España
Institución:Varias* (Consorci de Biblioteques Universitáries de Catalunya, Centre de Serveis Científics i Acadèmics de Catalunya)
Repositorio:Recercat. Dipósit de la Recerca de Catalunya
OAI Identifier:oai:recercat.cat:10230/70629
Acceso en línea:http://hdl.handle.net/10230/70629
http://dx.doi.org/10.1038/s41467-025-59430-w
Access Level:acceso abierto
Palabra clave:Biochemistry
Learning and memory
Molecular biology
Molecular neuroscience
RNA splicing
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network_name_str España
repository_id_str
dc.title.none.fl_str_mv A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
title A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
spellingShingle A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
Poliński, Patryk
Biochemistry
Learning and memory
Molecular biology
Molecular neuroscience
RNA splicing
title_short A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
title_full A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
title_fullStr A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
title_full_unstemmed A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
title_sort A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formation
dc.creator.none.fl_str_mv Poliński, Patryk
Miret-Cuesta, Marta
Zamora Moratalla, Alfonsa
Mantica, Federica
Cantero Recasens, Gerard, 1984-
Viana, Carlotta
Sabariego Navarro, Miguel
Normanno, Davide
Iñiguez, Luis P.
Bonnal, Sophie
Gómez-Riera, Raúl
Fanlo-Ucar, Hugo
Yap, Dominic S.
Martínez de Lagrán Cabredo, María
Fernández-Blanco, Álvaro
Rodríguez-Marin, Cristina
Permanyer, Jon
Fölsz, Orsolya
Domínguez-Sala, Eduardo
Sierra, Cesar
Wojnacki, José
Musoles Lleo, Juan Luis
Cosma, Maria Pia
Muñoz López, Francisco José, 1964-
Dierssen, Mara
Irimia Martínez, Manuel
author Poliński, Patryk
author_facet Poliński, Patryk
Miret-Cuesta, Marta
Zamora Moratalla, Alfonsa
Mantica, Federica
Cantero Recasens, Gerard, 1984-
Viana, Carlotta
Sabariego Navarro, Miguel
Normanno, Davide
Iñiguez, Luis P.
Bonnal, Sophie
Gómez-Riera, Raúl
Fanlo-Ucar, Hugo
Yap, Dominic S.
Martínez de Lagrán Cabredo, María
Fernández-Blanco, Álvaro
Rodríguez-Marin, Cristina
Permanyer, Jon
Fölsz, Orsolya
Domínguez-Sala, Eduardo
Sierra, Cesar
Wojnacki, José
Musoles Lleo, Juan Luis
Cosma, Maria Pia
Muñoz López, Francisco José, 1964-
Dierssen, Mara
Irimia Martínez, Manuel
author_role author
author2 Miret-Cuesta, Marta
Zamora Moratalla, Alfonsa
Mantica, Federica
Cantero Recasens, Gerard, 1984-
Viana, Carlotta
Sabariego Navarro, Miguel
Normanno, Davide
Iñiguez, Luis P.
Bonnal, Sophie
Gómez-Riera, Raúl
Fanlo-Ucar, Hugo
Yap, Dominic S.
Martínez de Lagrán Cabredo, María
Fernández-Blanco, Álvaro
Rodríguez-Marin, Cristina
Permanyer, Jon
Fölsz, Orsolya
Domínguez-Sala, Eduardo
Sierra, Cesar
Wojnacki, José
Musoles Lleo, Juan Luis
Cosma, Maria Pia
Muñoz López, Francisco José, 1964-
Dierssen, Mara
Irimia Martínez, Manuel
author2_role author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
dc.subject.none.fl_str_mv Biochemistry
Learning and memory
Molecular biology
Molecular neuroscience
RNA splicing
topic Biochemistry
Learning and memory
Molecular biology
Molecular neuroscience
RNA splicing
description Actin cytoskeleton dynamics is essential for proper nervous system development and function. A conserved set of neuronal-specific microexons influences multiple aspects of neurobiology; however, their roles in regulating the actin cytoskeleton are unknown. Here, we study a microexon in DAAM1, a formin-homology-2 (FH2) domain protein involved in actin reorganization. Microexon inclusion extends the linker region of the DAAM1 FH2 domain, altering actin polymerization. Genomic deletion of the microexon leads to neuritogenesis defects and increased calcium influx in differentiated neurons. Mice with this deletion exhibit postsynaptic defects, fewer immature dendritic spines, impaired long-term potentiation, and deficits in memory formation. These phenotypes are associated with increased RHOA/ROCK signaling, which regulates actin-cytoskeleton dynamics, and are partially rescued by treatment with a ROCK inhibitor. This study highlights the role of a conserved neuronal microexon in regulating actin dynamics and cognitive functioning.
publishDate 2025
dc.date.none.fl_str_mv 2025
2025
2025
dc.type.none.fl_str_mv info:eu-repo/semantics/article
info:eu-repo/semantics/publishedVersion
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv http://hdl.handle.net/10230/70629
http://dx.doi.org/10.1038/s41467-025-59430-w
http://hdl.handle.net/10230/70629
url http://hdl.handle.net/10230/70629
http://dx.doi.org/10.1038/s41467-025-59430-w
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
dc.relation.none.fl_str_mv Nat Commun. 2025 May 6;16(1):4210
info:eu-repo/grantAgreement/EC/H2020/101002275
info:eu-repo/grantAgreement/ES/2PE/PID2020-115040GB-I00
info:eu-repo/grantAgreement/ES/3PE/PID2022−141900OB-I00
info:eu-repo/grantAgreement/ES/2PE/PID2020-114080GB-I00
info:eu-repo/grantAgreement/ES/3PE/PID2023-1497670B-I00
info:eu-repo/grantAgreement/ES/3PE/PID2022-138245NB-I00
info:eu-repo/grantAgreement/EC/H2020/964342
info:eu-repo/grantAgreement/EC/H2020/721890
info:eu-repo/grantAgreement/EC/H2020/848077
dc.rights.none.fl_str_mv http://creativecommons.org/licenses/by-nc-nd/4.0/
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dc.publisher.none.fl_str_mv Nature Research
publisher.none.fl_str_mv Nature Research
dc.source.none.fl_str_mv reponame:Recercat. Dipósit de la Recerca de Catalunya
instname:Varias* (Consorci de Biblioteques Universitáries de Catalunya, Centre de Serveis Científics i Acadèmics de Catalunya)
instname_str Varias* (Consorci de Biblioteques Universitáries de Catalunya, Centre de Serveis Científics i Acadèmics de Catalunya)
reponame_str Recercat. Dipósit de la Recerca de Catalunya
collection Recercat. Dipósit de la Recerca de Catalunya
repository.name.fl_str_mv
repository.mail.fl_str_mv
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spelling A highly conserved neuronal microexon in DAAM1 controls actin dynamics, RHOA/ROCK signaling, and memory formationPoliński, PatrykMiret-Cuesta, MartaZamora Moratalla, AlfonsaMantica, FedericaCantero Recasens, Gerard, 1984-Viana, CarlottaSabariego Navarro, MiguelNormanno, DavideIñiguez, Luis P.Bonnal, SophieGómez-Riera, RaúlFanlo-Ucar, HugoYap, Dominic S.Martínez de Lagrán Cabredo, MaríaFernández-Blanco, ÁlvaroRodríguez-Marin, CristinaPermanyer, JonFölsz, OrsolyaDomínguez-Sala, EduardoSierra, CesarWojnacki, JoséMusoles Lleo, Juan LuisCosma, Maria PiaMuñoz López, Francisco José, 1964-Dierssen, MaraIrimia Martínez, ManuelBiochemistryLearning and memoryMolecular biologyMolecular neuroscienceRNA splicingActin cytoskeleton dynamics is essential for proper nervous system development and function. A conserved set of neuronal-specific microexons influences multiple aspects of neurobiology; however, their roles in regulating the actin cytoskeleton are unknown. Here, we study a microexon in DAAM1, a formin-homology-2 (FH2) domain protein involved in actin reorganization. Microexon inclusion extends the linker region of the DAAM1 FH2 domain, altering actin polymerization. Genomic deletion of the microexon leads to neuritogenesis defects and increased calcium influx in differentiated neurons. Mice with this deletion exhibit postsynaptic defects, fewer immature dendritic spines, impaired long-term potentiation, and deficits in memory formation. These phenotypes are associated with increased RHOA/ROCK signaling, which regulates actin-cytoskeleton dynamics, and are partially rescued by treatment with a ROCK inhibitor. This study highlights the role of a conserved neuronal microexon in regulating actin dynamics and cognitive functioning.The research has been funded by the European Research Council (ERC) under the European Union’s Horizon 2020 research and innovation program (ERCCoG-LS2-101002275 to M.I.), Spanish Ministry of Science and Innovation (PID2020-115040GB-I00 to M.I., PID2022−141900OB-I00 to M.D., PID2020- 114080GB-I00 to M.P.C., PID2023-1497670B-I00 to F.J.M. and PID2022-138245NB-I00 to E.H), the European Union’s Horizon 2020 research and innovation program under grant agreements No 964342 to M.P.C., 721890 to P.P. and 848077 to A.Z.M., the National Institutes of Health (NIH) (1R01NS137222-01 to M.D.) and AGAUR grants from Secretaria d’Universitats i Recerca del Departament d’Empresa iConeixement de la Generalitat de Catalunya to M.I., M.D. and M.P.C.. CRG acknowledges support of the Spanish Ministry of Science and Innovation through the Centro de Excelencia Severo Ochoa (CEX2020-001049-S, MCIN/AEI/10.13039/501100011033), and the Generalitat de Catalunya through the CERCA program. M.S.N. received an FPU fellowship (FPU19/04789) from Ministerio de Universidades. This research was in part supported by grants from the Simons Foundation and Canadian Institutes of Health Research to B.J.B., who also holds the University of Toronto Banbury Chair in Medical Research and the Canada Research Chair in RNA Biology and Genomics.Nature Research202520252025info:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionapplication/pdfapplication/pdfhttp://hdl.handle.net/10230/70629http://dx.doi.org/10.1038/s41467-025-59430-whttp://hdl.handle.net/10230/70629reponame:Recercat. Dipósit de la Recerca de Catalunyainstname:Varias* (Consorci de Biblioteques Universitáries de Catalunya, Centre de Serveis Científics i Acadèmics de Catalunya)InglésNat Commun. 2025 May 6;16(1):4210info:eu-repo/grantAgreement/EC/H2020/101002275info:eu-repo/grantAgreement/ES/2PE/PID2020-115040GB-I00info:eu-repo/grantAgreement/ES/3PE/PID2022−141900OB-I00info:eu-repo/grantAgreement/ES/2PE/PID2020-114080GB-I00info:eu-repo/grantAgreement/ES/3PE/PID2023-1497670B-I00info:eu-repo/grantAgreement/ES/3PE/PID2022-138245NB-I00info:eu-repo/grantAgreement/EC/H2020/964342info:eu-repo/grantAgreement/EC/H2020/721890info:eu-repo/grantAgreement/EC/H2020/848077© The Author(s) 2025. Open Access This article is licensed under a Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 International License, which permits any non-commercial use, sharing, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if you modified the licensed material. You do not have permission under this licence to share adapted material derived from this article or parts of it. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by-nc-nd/4.0/.http://creativecommons.org/licenses/by-nc-nd/4.0/info:eu-repo/semantics/openAccessoai:recercat.cat:10230/706292026-05-29T05:05:01Z
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