Transportome remodeling of a symbiotic microalga inside a planktonic host

Metabolic exchange is one of the foundations of symbiotic associations between organisms and is a driving force in evolution. In the ocean, photosymbiosis between heterotrophic hosts and microalgae is powered by photosynthesis and relies on the transfer of organic carbon to the host (e.g. sugars). Y...

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Autores: Juéry, Caroline, Auladell, Adria, Füssy, Zoltan, Chevalier, Fabien, Yee, Daniel P., Pelletier, Eric, Corre, Erwan, Allen, Andrew E., Richter, Daniel J., Decelle, Johan
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2024
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/383328
Acceso en línea:http://hdl.handle.net/10261/383328
https://api.elsevier.com/content/abstract/scopus_id/85214553157
Access Level:acceso abierto
Palabra clave:Transporters
Carbon exchange
Metatranscriptomics
Microalga
Planktonic photosymbiosis
Protists
Single-cell transcriptomic
Sugars
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dc.title.none.fl_str_mv Transportome remodeling of a symbiotic microalga inside a planktonic host
title Transportome remodeling of a symbiotic microalga inside a planktonic host
spellingShingle Transportome remodeling of a symbiotic microalga inside a planktonic host
Juéry, Caroline
Transporters
Carbon exchange
Metatranscriptomics
Microalga
Planktonic photosymbiosis
Protists
Single-cell transcriptomic
Sugars
title_short Transportome remodeling of a symbiotic microalga inside a planktonic host
title_full Transportome remodeling of a symbiotic microalga inside a planktonic host
title_fullStr Transportome remodeling of a symbiotic microalga inside a planktonic host
title_full_unstemmed Transportome remodeling of a symbiotic microalga inside a planktonic host
title_sort Transportome remodeling of a symbiotic microalga inside a planktonic host
dc.creator.none.fl_str_mv Juéry, Caroline
Auladell, Adria
Füssy, Zoltan
Chevalier, Fabien
Yee, Daniel P.
Pelletier, Eric
Corre, Erwan
Allen, Andrew E.
Richter, Daniel J.
Decelle, Johan
author Juéry, Caroline
author_facet Juéry, Caroline
Auladell, Adria
Füssy, Zoltan
Chevalier, Fabien
Yee, Daniel P.
Pelletier, Eric
Corre, Erwan
Allen, Andrew E.
Richter, Daniel J.
Decelle, Johan
author_role author
author2 Auladell, Adria
Füssy, Zoltan
Chevalier, Fabien
Yee, Daniel P.
Pelletier, Eric
Corre, Erwan
Allen, Andrew E.
Richter, Daniel J.
Decelle, Johan
author2_role author
author
author
author
author
author
author
author
author
dc.contributor.none.fl_str_mv Centre National de la Recherche Scientifique (France)
European Commission
European Research Council
National Oceanic and Atmospheric Administration (US)
National Science Foundation (US)
Department of Energy (US)
Generalitat de Catalunya
Consejo Superior de Investigaciones Científicas [https://ror.org/02gfc7t72]
dc.subject.none.fl_str_mv Transporters
Carbon exchange
Metatranscriptomics
Microalga
Planktonic photosymbiosis
Protists
Single-cell transcriptomic
Sugars
topic Transporters
Carbon exchange
Metatranscriptomics
Microalga
Planktonic photosymbiosis
Protists
Single-cell transcriptomic
Sugars
description Metabolic exchange is one of the foundations of symbiotic associations between organisms and is a driving force in evolution. In the ocean, photosymbiosis between heterotrophic hosts and microalgae is powered by photosynthesis and relies on the transfer of organic carbon to the host (e.g. sugars). Yet, the identity of transferred carbohydrates as well as the molecular mechanisms that drive this exchange remain largely unknown, especially in unicellular photosymbioses that are widespread in the open ocean. Combining genomics, single-holobiont transcriptomics, and environmental metatranscriptomics, we revealed the transportome of the marine microalga Phaeocystis in symbiosis within acantharia, with a focus on sugar transporters. At the genomic level, the sugar transportome of Phaeocystis is comparable to non-symbiotic haptophytes. By contrast, we found significant remodeling of the expression of the transportome in symbiotic microalgae compared to the free-living stage. More particularly, 36% of sugar transporter genes were differentially expressed. Several of them, such as GLUTs, TPTs, and aquaporins, with glucose, triose-phosphate sugars, and glycerol as potential substrates, were upregulated at the holobiont and community level. We also showed that algal sugar transporter genes exhibit distinct temporal expression patterns during the day. This reprogramed transportome indicates that symbiosis has a major impact on sugar fluxes within and outside the algal cell, and highlights the complexity and the dynamics of metabolic exchanges between partners. This study improves our understanding of the molecular players of the metabolic connectivity underlying the ecological success of planktonic photosymbiosis and paves the way for more studies on transporters across photosymbiotic models.
publishDate 2024
dc.date.none.fl_str_mv 2024
2025
2025
dc.type.none.fl_str_mv info:eu-repo/semantics/article
http://purl.org/coar/resource_type/c_6501
Publisher's version
info:eu-repo/semantics/publishedVersion
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv http://hdl.handle.net/10261/383328
https://api.elsevier.com/content/abstract/scopus_id/85214553157
url http://hdl.handle.net/10261/383328
https://api.elsevier.com/content/abstract/scopus_id/85214553157
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
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The underlying dataset has been published as supplementary material of the article in the publisher platform at DOI https://doi.org/10.1093/ismejo/wrae239
https://doi.org/10.1093/ismejo/wrae239

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dc.publisher.none.fl_str_mv Oxford University Press
publisher.none.fl_str_mv Oxford University Press
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spelling Transportome remodeling of a symbiotic microalga inside a planktonic hostJuéry, CarolineAuladell, AdriaFüssy, ZoltanChevalier, FabienYee, Daniel P.Pelletier, EricCorre, ErwanAllen, Andrew E.Richter, Daniel J.Decelle, JohanTransportersCarbon exchangeMetatranscriptomicsMicroalgaPlanktonic photosymbiosisProtistsSingle-cell transcriptomicSugarsMetabolic exchange is one of the foundations of symbiotic associations between organisms and is a driving force in evolution. In the ocean, photosymbiosis between heterotrophic hosts and microalgae is powered by photosynthesis and relies on the transfer of organic carbon to the host (e.g. sugars). Yet, the identity of transferred carbohydrates as well as the molecular mechanisms that drive this exchange remain largely unknown, especially in unicellular photosymbioses that are widespread in the open ocean. Combining genomics, single-holobiont transcriptomics, and environmental metatranscriptomics, we revealed the transportome of the marine microalga Phaeocystis in symbiosis within acantharia, with a focus on sugar transporters. At the genomic level, the sugar transportome of Phaeocystis is comparable to non-symbiotic haptophytes. By contrast, we found significant remodeling of the expression of the transportome in symbiotic microalgae compared to the free-living stage. More particularly, 36% of sugar transporter genes were differentially expressed. Several of them, such as GLUTs, TPTs, and aquaporins, with glucose, triose-phosphate sugars, and glycerol as potential substrates, were upregulated at the holobiont and community level. We also showed that algal sugar transporter genes exhibit distinct temporal expression patterns during the day. This reprogramed transportome indicates that symbiosis has a major impact on sugar fluxes within and outside the algal cell, and highlights the complexity and the dynamics of metabolic exchanges between partners. This study improves our understanding of the molecular players of the metabolic connectivity underlying the ecological success of planktonic photosymbiosis and paves the way for more studies on transporters across photosymbiotic models.Research was supported by CNRS and ATIP-Avenir program funding, and by the European Union (BIOcean5D—GA#101059915 and ERC—SymbiOCEAN—101088661). This project has received funding from the European Union’s Horizon 2020 research and innovation program under grant agreement no. 824110 for the sequencing (EASI Genomics). This project has received funding from the European Union’s Horizon 2020 research and innovation program under grant agreement no. 824110 for the sequencing (EASI Genomics). Z.F. and A.A. were funded by National Oceanic and Atmospheric Administration grant NA19NOS4780181 (to A.E.A.), the National Science Foundation (NSF-OCE-1756884 to A.E.A.), and the Simons Collaboration on Principles of Microbial Ecosystems (PriME; grant ID: 970820 to A.E.A.). Their work (proposal: 10.46936/10.25585/60001426) conducted in the US Department of Energy Joint Genome Institute (https://ror.org/04xm1d337), a DOE Office of Science User Facility, is supported by the Office of Science of the US Department of Energy operated under contract no. DE-AC02-05CH11231. A.A. and D.J.R. received funding from the European Research Council (ERC) under the European Union’s Horizon 2020 research and innovation program (grant agreement 949745—GROWCEAN—ERC-2020-STG) and from the Departament de Recerca i Universitats de la Generalitat de Catalunya (exp. 2021 SGR 00751).Peer reviewedOxford University PressCentre National de la Recherche Scientifique (France)European CommissionEuropean Research CouncilNational Oceanic and Atmospheric Administration (US)National Science Foundation (US)Department of Energy (US)Generalitat de CatalunyaConsejo Superior de Investigaciones Científicas [https://ror.org/02gfc7t72]202520252024info:eu-repo/semantics/articlehttp://purl.org/coar/resource_type/c_6501Publisher's versioninfo:eu-repo/semantics/publishedVersionapplication/pdfhttp://hdl.handle.net/10261/383328https://api.elsevier.com/content/abstract/scopus_id/85214553157reponame:DIGITAL.CSIC. Repositorio Institucional del CSICinstname:Consejo Superior de Investigaciones Científicas (CSIC)Inglés#PLACEHOLDER_PARENT_METADATA_VALUE##PLACEHOLDER_PARENT_METADATA_VALUE##PLACEHOLDER_PARENT_METADATA_VALUE##PLACEHOLDER_PARENT_METADATA_VALUE#info:eu-repo/grantAgreement/EC/HE/101059915info:eu-repo/grantAgreement/EC/HE/101088661info:eu-repo/grantAgreement/EC/H2020/824110info:eu-repo/grantAgreement/EC/H2020/949745The underlying dataset has been published as supplementary material of the article in the publisher platform at DOI https://doi.org/10.1093/ismejo/wrae239https://doi.org/10.1093/ismejo/wrae239Síinfo:eu-repo/semantics/openAccessoai:digital.csic.es:10261/3833282026-05-22T06:33:51Z
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