The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium

[EN] The four NGATHA genes (NGA) form a small subfamily within the large family of B3-domain transcription factors of Arabidopsis thaliana. NGA genes act redundantly to direct the development of the apical tissues of the gynoecium, the style, and the stigma. Previous studies indicate that NGA genes...

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Autores: Martínez Fernández, Irene, Sanchis, Sofía, Marini, Naciele, Navarrete Gomez, Maria Luisa, Oliveira, Antonio C., Colombo, Lucia, Balanzà, Vicente|||0000-0002-1761-1524, Ballester Fuentes, Patricia, FERRANDIZ MAESTRE, CRISTINA|||0000-0002-2460-1068
Tipo de documento: artigo
Data de publicação:2014
País:España
Recursos:Universitat Politècnica de València (UPV)
Repositório:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
Idioma:inglês
OAI Identifier:oai:riunet.upv.es:10251/67957
Acesso em linha:https://riunet.upv.es/handle/10251/67957
Access Level:Acceso aberto
Palavra-chave:Gynoecium development
Ngatha
Auxin synthesis
Auxin transport
Auxin response factors
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spelling The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoeciumMartínez Fernández, IreneSanchis, SofíaMarini, NacieleNavarrete Gomez, Maria LuisaOliveira, Antonio C.Colombo, LuciaBalanzà, Vicente|||0000-0002-1761-1524Ballester Fuentes, PatriciaFERRANDIZ MAESTRE, CRISTINA|||0000-0002-2460-1068Gynoecium developmentNgathaAuxin synthesisAuxin transportAuxin response factors[EN] The four NGATHA genes (NGA) form a small subfamily within the large family of B3-domain transcription factors of Arabidopsis thaliana. NGA genes act redundantly to direct the development of the apical tissues of the gynoecium, the style, and the stigma. Previous studies indicate that NGA genes could exert this function at least partially by directing the synthesis of auxin at the distal end of the developing gynoecium through the upregulation of two different YUCCA genes, which encode flavin monooxygenases involved in auxin biosynthesis. We have compared three developing pistil transcriptome data sets from wildtype, nga quadruple mutants, and a 35S::NGA3 line. The differentially expressed genes showed a significant enrichment for auxin-related genes, supporting the idea of NGA genes as major regulators of auxin accumulation and distribution within the developing gynoecium. We have introduced reporter lines for several of these differentially expressed genes involved in synthesis, transport and response to auxin in NGA gain- and loss-of-function backgrounds. We present here a detailed map of the response of these reporters to NGA misregulation that could help to clarify the role of NGA in auxin-mediated gynoecium morphogenesis. Our data point to a very reduced auxin synthesis in the developing apical gynoecium of nga mutants, likely responsible for the lack of DR5rev::GFP reporter activity observed in these mutants. In addition, NGA altered activity affects the expression of protein kinases that regulate the cellular localization of auxin efflux regulators, and thus likely impact auxin transport. Finally, protein accumulation in pistils of several ARFs was differentially affected by nga mutations or NGA overexpression, suggesting that these accumulation patterns depend not only on auxin distribution but could be also regulated by transcriptional networks involving NGA factors.This work was supported by the Spanish Ministerio de Ciencia e Innovacion (grant no. BIO2009–09920 to Cristina Ferrándiz), the Fondazione Cariplo (grant “Fruitalia” to Lucia Colombo and Cristina Ferrándiz), and the European Union (grant no. FP7–PEOPLE–PIRSES–2009–247589 to Lucia Colombo, Cristina Ferrándiz, Antonio C. Oliveira).Frontiers MediaInstituto Universitario Mixto de Biología Molecular y Celular de PlantasEuropean CommissionMinisterio de Ciencia e InnovaciónFondazione CariploRepositorio Institucional de la Universitat Politècnica de València Riunet20142014-05-21journal articlehttp://purl.org/coar/resource_type/c_6501VoRhttp://purl.org/coar/version/c_970fb48d4fbd8a85info:eu-repo/semantics/articleapplication/pdfhttps://riunet.upv.es/handle/10251/67957reponame:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valénciainstname:Universitat Politècnica de València (UPV)InglésengEuropean Commission https://doi.org/10.13039/501100000780 FP7 247589Ministerio de Ciencia e Innovación http://dx.doi.org/10.13039/501100004837 BIO2009-09920 Conservacion De Los Modelos Geneticos Que Explican La Morfogenesis Del Carpelo Y El Fruto En Las Eudicotiledoneasopen accesshttp://purl.org/coar/access_right/c_abf2Reserva de todos los derechoshttp://rightsstatements.org/vocab/InC/1.0/info:eu-repo/semantics/openAccessoai:riunet.upv.es:10251/679572026-06-13T07:49:27Z
dc.title.none.fl_str_mv The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
title The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
spellingShingle The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
Martínez Fernández, Irene
Gynoecium development
Ngatha
Auxin synthesis
Auxin transport
Auxin response factors
title_short The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
title_full The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
title_fullStr The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
title_full_unstemmed The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
title_sort The effect of NGATHA altered activity on auxin signaling pathways within the Arabidopsis gynoecium
dc.creator.none.fl_str_mv Martínez Fernández, Irene
Sanchis, Sofía
Marini, Naciele
Navarrete Gomez, Maria Luisa
Oliveira, Antonio C.
Colombo, Lucia
Balanzà, Vicente|||0000-0002-1761-1524
Ballester Fuentes, Patricia
FERRANDIZ MAESTRE, CRISTINA|||0000-0002-2460-1068
author Martínez Fernández, Irene
author_facet Martínez Fernández, Irene
Sanchis, Sofía
Marini, Naciele
Navarrete Gomez, Maria Luisa
Oliveira, Antonio C.
Colombo, Lucia
Balanzà, Vicente|||0000-0002-1761-1524
Ballester Fuentes, Patricia
FERRANDIZ MAESTRE, CRISTINA|||0000-0002-2460-1068
author_role author
author2 Sanchis, Sofía
Marini, Naciele
Navarrete Gomez, Maria Luisa
Oliveira, Antonio C.
Colombo, Lucia
Balanzà, Vicente|||0000-0002-1761-1524
Ballester Fuentes, Patricia
FERRANDIZ MAESTRE, CRISTINA|||0000-0002-2460-1068
author2_role author
author
author
author
author
author
author
author
dc.contributor.none.fl_str_mv Instituto Universitario Mixto de Biología Molecular y Celular de Plantas
European Commission
Ministerio de Ciencia e Innovación
Fondazione Cariplo
Repositorio Institucional de la Universitat Politècnica de València Riunet
dc.subject.none.fl_str_mv Gynoecium development
Ngatha
Auxin synthesis
Auxin transport
Auxin response factors
topic Gynoecium development
Ngatha
Auxin synthesis
Auxin transport
Auxin response factors
description [EN] The four NGATHA genes (NGA) form a small subfamily within the large family of B3-domain transcription factors of Arabidopsis thaliana. NGA genes act redundantly to direct the development of the apical tissues of the gynoecium, the style, and the stigma. Previous studies indicate that NGA genes could exert this function at least partially by directing the synthesis of auxin at the distal end of the developing gynoecium through the upregulation of two different YUCCA genes, which encode flavin monooxygenases involved in auxin biosynthesis. We have compared three developing pistil transcriptome data sets from wildtype, nga quadruple mutants, and a 35S::NGA3 line. The differentially expressed genes showed a significant enrichment for auxin-related genes, supporting the idea of NGA genes as major regulators of auxin accumulation and distribution within the developing gynoecium. We have introduced reporter lines for several of these differentially expressed genes involved in synthesis, transport and response to auxin in NGA gain- and loss-of-function backgrounds. We present here a detailed map of the response of these reporters to NGA misregulation that could help to clarify the role of NGA in auxin-mediated gynoecium morphogenesis. Our data point to a very reduced auxin synthesis in the developing apical gynoecium of nga mutants, likely responsible for the lack of DR5rev::GFP reporter activity observed in these mutants. In addition, NGA altered activity affects the expression of protein kinases that regulate the cellular localization of auxin efflux regulators, and thus likely impact auxin transport. Finally, protein accumulation in pistils of several ARFs was differentially affected by nga mutations or NGA overexpression, suggesting that these accumulation patterns depend not only on auxin distribution but could be also regulated by transcriptional networks involving NGA factors.
publishDate 2014
dc.date.none.fl_str_mv 2014
2014-05-21
dc.type.none.fl_str_mv journal article
http://purl.org/coar/resource_type/c_6501
VoR
http://purl.org/coar/version/c_970fb48d4fbd8a85
dc.type.openaire.fl_str_mv info:eu-repo/semantics/article
format article
dc.identifier.none.fl_str_mv https://riunet.upv.es/handle/10251/67957
url https://riunet.upv.es/handle/10251/67957
dc.language.none.fl_str_mv Inglés
eng
language_invalid_str_mv Inglés
language eng
dc.relation.none.fl_str_mv European Commission https://doi.org/10.13039/501100000780 FP7 247589
Ministerio de Ciencia e Innovación http://dx.doi.org/10.13039/501100004837 BIO2009-09920 Conservacion De Los Modelos Geneticos Que Explican La Morfogenesis Del Carpelo Y El Fruto En Las Eudicotiledoneas
dc.rights.none.fl_str_mv open access
http://purl.org/coar/access_right/c_abf2
Reserva de todos los derechos
http://rightsstatements.org/vocab/InC/1.0/
dc.rights.openaire.fl_str_mv info:eu-repo/semantics/openAccess
rights_invalid_str_mv open access
http://purl.org/coar/access_right/c_abf2
Reserva de todos los derechos
http://rightsstatements.org/vocab/InC/1.0/
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
dc.publisher.none.fl_str_mv Frontiers Media
publisher.none.fl_str_mv Frontiers Media
dc.source.none.fl_str_mv reponame:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
instname:Universitat Politècnica de València (UPV)
instname_str Universitat Politècnica de València (UPV)
reponame_str RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
collection RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
repository.name.fl_str_mv
repository.mail.fl_str_mv
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