Caveolin-1 dolines form a distinct and rapid caveolae-independent mechanoadaptation system

In response to diferent types and intensities of mechanical force, cells modulate their physical properties and adapt their plasma membrane (PM). Caveolae are PM nano-invaginations that contribute to mechanoadaptation, bufering tension changes. However, whether core caveolar proteins contribute to P...

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Detalles Bibliográficos
Autores: Lolo, Fidel Nicolás, Walani, Nikhil, Seemann, Eric, Zalvidea, Dobryna, Pavón, Dácil María, Cojoc, Gheorghe, Zamai, Moreno, Viaris de Lesegno, Christine, Martínez de Benito, Fernando, Sánchez Álvarez, Miguel, Uriarte, Juan José, Echarri, Asier, Jiménez Carretero, Daniel, Escolano, Joan Carles, Sánchez Donoso, Susana Angélica, Arroyo Balaguer, Marino|||0000-0003-1647-940X
Tipo de recurso: artículo
Fecha de publicación:2022
País:España
Institución:Universitat Politècnica de Catalunya (UPC)
Repositorio:UPCommons. Portal del coneixement obert de la UPC
Idioma:inglés
OAI Identifier:oai:upcommons.upc.edu:2117/386804
Acceso en línea:https://hdl.handle.net/2117/386804
https://dx.doi.org/10.1038/s41556-022-01034-3
Access Level:acceso abierto
Palabra clave:Biology
Biomathematics
Biologia
Biomatemàtica
Classificació AMS::92 Biology and other natural sciences::92C Physiological, cellular and medical topics
Classificació AMS::92 Biology and other natural sciences::92B Mathematical biology in general
Àrees temàtiques de la UPC::Matemàtiques i estadística::Matemàtica aplicada a les ciències
Descripción
Sumario:In response to diferent types and intensities of mechanical force, cells modulate their physical properties and adapt their plasma membrane (PM). Caveolae are PM nano-invaginations that contribute to mechanoadaptation, bufering tension changes. However, whether core caveolar proteins contribute to PM tension accommodation independently from the caveolar assembly is unknown. Here we provide experimental and computational evidence supporting that caveolin-1 confers deformability and mechanoprotection independently from caveolae, through modulation of PM curvature. Freeze-fracture electron microscopy reveals that caveolin-1 stabilizes non-caveolar invaginations—dolines—capable of responding to low-medium mechanical forces, impacting downstream mechanotransduction and conferring mechanoprotection to cells devoid of caveolae. Upon cavin-1/PTRF binding, doline size is restricted and membrane bufering is limited to relatively high forces, capable of fattening caveolae. Thus, caveolae and dolines constitute two distinct albeit complementary components of a bufering system that allows cells to adapt efciently to a broad range of mechanical stimuli.