Functional analysis of two novel retinitis pigmentosa mutations reveals structural role of the third transmembrane helix in rhodopsin photoactivation

Mutations in the visual receptor rhodopsin are a common cause of inherited retinal diseases. Many of these mutations occur within the transmembrane core of the receptor and can impact in its folding and function. In this study, we investigate two mutations associated with the retinal degenerative di...

Descripción completa

Detalles Bibliográficos
Autores: Fernández González, Pol|||0000-0001-6136-763X, Cruz Sáez, Alejandro|||0000-0003-0928-0718, Wang, Feifei, Cordomí, Arnau, Morillo Cazorla, Margarita|||0000-0003-3624-9900, Pérez González, Juan Jesús|||0000-0002-0748-8147, Garriga Solé, Pere|||0000-0003-4234-8382
Tipo de recurso: artículo
Fecha de publicación:2026
País:España
Institución:Universitat Politècnica de Catalunya (UPC)
Repositorio:UPCommons. Portal del coneixement obert de la UPC
Idioma:inglés
OAI Identifier:oai:dnet:upcommonspor::e8b9748fd4f3f750ca2f55d4f99fdc71
Acceso en línea:https://hdl.handle.net/2117/461064
https://dx.doi.org/10.1016/j.ijbiomac.2026.152176
Access Level:acceso abierto
Palabra clave:Rhodopsin
Visual phototransduction
Protein misfolding
Retinal degeneration
Àrees temàtiques de la UPC::Enginyeria química
Descripción
Sumario:Mutations in the visual receptor rhodopsin are a common cause of inherited retinal diseases. Many of these mutations occur within the transmembrane core of the receptor and can impact in its folding and function. In this study, we investigate two mutations associated with the retinal degenerative disease retinitis pigmentosa, T1083.23P and G1213.36R, both located in the third transmembrane helix of rhodopsin and in proximity to key structural residues essential for rhodopsin stability and proper folding. We find that the G1213.36R mutant fails to regenerate with 11-cis-retinal, suggesting that the mutation severely impairs chromophore binding and proper folding. In contrast, the T1083.23P mutant regenerates efficiently, shows normal photobleaching and acidification profiles, and similar chromophore regeneration to that of the wild-type protein. However, the T1083.23P mutation leads to reduced thermal and chemical stability of its dark state, and a delayed formation of the active conformation. These conformational alterations correlate with a slower and less efficient activation of transducin, indicating that T1083.23P partially decouples the light-induced response from downstream G-protein signaling. Together, these findings demonstrate that amino acid substitutions in the third transmembrane helix of rhodopsin can have diverse molecular consequences: from the severe impairment of chromophore binding in G1213.36R to selective destabilization and signaling defects in the T1083.23P case. Our results emphasize the importance of the third transmembrane helix in supporting the conformational changes necessary for rhodopsin activation. and contribute to a deeper understanding of how specific mutations in rhodopsin may elicit distinct pathogenic mechanisms underlying retinal degeneration