Cyanobacterial Septal Junctions: Properties and Regulation

Heterocyst-forming cyanobacteria are multicellular organisms that grow as chains of cells (filaments or trichomes) in which the cells exchange regulators and nutrients. In this article, we review the morphological, physiological and genetic data that have led to our current understanding of intercel...

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Detalles Bibliográficos
Autores: Flores, Enrique, Nieves-Morión, Mercedes, Mullineaux, Conrad W.
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2018
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/178923
Acceso en línea:http://hdl.handle.net/10261/178923
Access Level:acceso abierto
Palabra clave:Anabaena
Bacterial multicellularity
Intercellular communication
Descripción
Sumario:Heterocyst-forming cyanobacteria are multicellular organisms that grow as chains of cells (filaments or trichomes) in which the cells exchange regulators and nutrients. In this article, we review the morphological, physiological and genetic data that have led to our current understanding of intercellular communication in these organisms. Intercellular molecular exchange appears to take place by simple diffusion through proteinaceous structures, known as septal junctions, which connect the adjacent cells in the filament and traverse the septal peptidoglycan through perforations known as nanopores. Proteins that are necessary to produce, and that may be components of, the septal junctions―SepJ, FraC and FraD―have been identified in the heterocyst-forming cyanobacterium <i>Anabaena</i> sp. strain PCC 7120 model. Additionally, several proteins that are necessary to produce a normal number of nanopores and functional septal junctions have been identified, including AmiC-type amidases, peptidoglycan-binding proteins and some membrane transporters. Available reports and reevaluation of intercellular molecular transfer data for some mutants of <i>Anabaena</i> suggest that the septal junctions can be regulated, likely by a mechanism of gating.