HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation

[EN] The gynoecium is critical for the reproduction of flowering plants as it contains the ovules and the tissues that foster pollen germination, growth, and guidance. These tissues, known as the reproductive tract (ReT), comprise the stigma, style, and transmitting tract (TT). The ReT and ovules or...

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Autores: Manrique-Urpí, Silvia|||0000-0001-6241-4143, Bombarely A, Cavalleri, Alex, Guazzotti, Andrea, Villarino, Gonzalo H., Simonini, Sara, Higashiyama, Tetsuya, Grossniklaus, Ueli, Mizzotti, Chiara, Pereira, Ana Marta, Coimbra, Silvia, Sankaranarayanan, Subramanian, Onelli, Elisabetta, Masiero, Simona, Franks, Robert G.
Tipo de recurso: artículo
Fecha de publicación:2024
País:España
Institución:Universitat Politècnica de València (UPV)
Repositorio:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
Idioma:inglés
OAI Identifier:oai:dnet:riunet______::6b0a516ebfdf8c96bc387baf54e0ede3
Acceso en línea:https://riunet.upv.es/handle/10251/234136
Access Level:acceso abierto
Palabra clave:Gynoecium development
Arabidopsis thaliana
Carpel margin meristem
Histone deacetylase 19
SHOOT MERISTEMLESS
Ovule formation
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dc.title.none.fl_str_mv HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
title HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
spellingShingle HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
Manrique-Urpí, Silvia|||0000-0001-6241-4143
Gynoecium development
Arabidopsis thaliana
Carpel margin meristem
Histone deacetylase 19
SHOOT MERISTEMLESS
Ovule formation
title_short HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
title_full HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
title_fullStr HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
title_full_unstemmed HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
title_sort HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract Differentiation
dc.creator.none.fl_str_mv Manrique-Urpí, Silvia|||0000-0001-6241-4143
Bombarely A
Cavalleri, Alex
Guazzotti, Andrea
Villarino, Gonzalo H.
Simonini, Sara
Higashiyama, Tetsuya
Grossniklaus, Ueli
Mizzotti, Chiara
Pereira, Ana Marta
Coimbra, Silvia
Sankaranarayanan, Subramanian
Onelli, Elisabetta
Masiero, Simona
Franks, Robert G.
author Manrique-Urpí, Silvia|||0000-0001-6241-4143
author_facet Manrique-Urpí, Silvia|||0000-0001-6241-4143
Bombarely A
Cavalleri, Alex
Guazzotti, Andrea
Villarino, Gonzalo H.
Simonini, Sara
Higashiyama, Tetsuya
Grossniklaus, Ueli
Mizzotti, Chiara
Pereira, Ana Marta
Coimbra, Silvia
Sankaranarayanan, Subramanian
Onelli, Elisabetta
Masiero, Simona
Franks, Robert G.
author_role author
author2 Bombarely A
Cavalleri, Alex
Guazzotti, Andrea
Villarino, Gonzalo H.
Simonini, Sara
Higashiyama, Tetsuya
Grossniklaus, Ueli
Mizzotti, Chiara
Pereira, Ana Marta
Coimbra, Silvia
Sankaranarayanan, Subramanian
Onelli, Elisabetta
Masiero, Simona
Franks, Robert G.
author2_role author
author
author
author
author
author
author
author
author
author
author
author
author
author
dc.contributor.none.fl_str_mv Departamento de Biotecnología
Instituto Universitario de Conservación y Mejora de la Agrodiversidad Valenciana
Escuela Técnica Superior de Ingeniería Agronómica y del Medio Natural
European Commission
Swiss National Science Foundation
European Molecular Biology Organization
Fundação para a Ciência e a Tecnologia, Portugal
Ministero dell'Istruzione, dell'Università e della Ricerca
Repositorio Institucional de la Universitat Politècnica de València Riunet
dc.subject.none.fl_str_mv Gynoecium development
Arabidopsis thaliana
Carpel margin meristem
Histone deacetylase 19
SHOOT MERISTEMLESS
Ovule formation
topic Gynoecium development
Arabidopsis thaliana
Carpel margin meristem
Histone deacetylase 19
SHOOT MERISTEMLESS
Ovule formation
description [EN] The gynoecium is critical for the reproduction of flowering plants as it contains the ovules and the tissues that foster pollen germination, growth, and guidance. These tissues, known as the reproductive tract (ReT), comprise the stigma, style, and transmitting tract (TT). The ReT and ovules originate from the carpel margin meristem (CMM) within the pistil. SHOOT MERISTEMLESS (STM) is a key transcription factor for meristem formation and maintenance. In all above-ground meristems, including the CMM, local STM downregulation is required for organ formation. However, how this downregulation is achieved in the CMM is unknown. Here, we have studied the role of HISTONE DEACETYLASE 19 (HDA19) in Arabidopsis (Arabidopsis thaliana) during ovule and ReT differentiation based on the observation that the hda19-3 mutant displays a reduced ovule number and fails to differentiate the TT properly. Fluorescence-activated cell sorting coupled with RNA-sequencing revealed that in the CMM of hda19-3 mutants, genes promoting organ development are downregulated while meristematic markers, including STM, are upregulated. HDA19 was essential to downregulate STM in the CMM, thereby allowing ovule formation and TT differentiation. STM is ectopically expressed in hda19-3 at intermediate stages of pistil development, and its downregulation by RNA interference alleviated the hda19-3 phenotype. Chromatin immunoprecipitation assays indicated that STM is a direct target of HDA19 during pistil development and that the transcription factor SEEDSTICK is also required to regulate STM via histone acetylation. Thus, we identified factors required for the downregulation of STM in the CMM, which is necessary for organogenesis and tissue differentiation.
publishDate 2024
dc.date.none.fl_str_mv 2024
2024-01-01
dc.type.none.fl_str_mv journal article
http://purl.org/coar/resource_type/c_6501
VoR
http://purl.org/coar/version/c_970fb48d4fbd8a85
dc.type.openaire.fl_str_mv info:eu-repo/semantics/article
format article
dc.identifier.none.fl_str_mv https://riunet.upv.es/handle/10251/234136
url https://riunet.upv.es/handle/10251/234136
dc.language.none.fl_str_mv Inglés
eng
language_invalid_str_mv Inglés
language eng
dc.relation.none.fl_str_mv European Commission https://doi.org/10.13039/501100000780 FP7 612640 The physiology and genetics of fruit formation: from genes to networks
European Commission https://doi.org/10.13039/501100000780 H2020 690946 Sexual Plant Reproduction ¿ Seed formation
Fundação para a Ciência e a Tecnologia, Portugal https://doi.org/10.13039/501100001871 POCI-01-0145-FEDER-027839
Swiss National Science Foundation https://doi.org/10.13039/501100001711 31003A_179553
dc.rights.none.fl_str_mv open access
http://purl.org/coar/access_right/c_abf2
Reconocimiento (by)
http://creativecommons.org/licenses/by/4.0/
dc.rights.openaire.fl_str_mv info:eu-repo/semantics/openAccess
rights_invalid_str_mv open access
http://purl.org/coar/access_right/c_abf2
Reconocimiento (by)
http://creativecommons.org/licenses/by/4.0/
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
dc.publisher.none.fl_str_mv American Society of Plant Physiologists
publisher.none.fl_str_mv American Society of Plant Physiologists
dc.source.none.fl_str_mv reponame:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
instname:Universitat Politècnica de València (UPV)
instname_str Universitat Politècnica de València (UPV)
reponame_str RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
collection RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
repository.name.fl_str_mv
repository.mail.fl_str_mv
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spelling HISTONE DEACETYLASE19 Controls Ovule Number Determination and Transmitting Tract DifferentiationManrique-Urpí, Silvia|||0000-0001-6241-4143Bombarely ACavalleri, AlexGuazzotti, AndreaVillarino, Gonzalo H.Simonini, SaraHigashiyama, TetsuyaGrossniklaus, UeliMizzotti, ChiaraPereira, Ana MartaCoimbra, SilviaSankaranarayanan, SubramanianOnelli, ElisabettaMasiero, SimonaFranks, Robert G.Gynoecium developmentArabidopsis thalianaCarpel margin meristemHistone deacetylase 19SHOOT MERISTEMLESSOvule formation[EN] The gynoecium is critical for the reproduction of flowering plants as it contains the ovules and the tissues that foster pollen germination, growth, and guidance. These tissues, known as the reproductive tract (ReT), comprise the stigma, style, and transmitting tract (TT). The ReT and ovules originate from the carpel margin meristem (CMM) within the pistil. SHOOT MERISTEMLESS (STM) is a key transcription factor for meristem formation and maintenance. In all above-ground meristems, including the CMM, local STM downregulation is required for organ formation. However, how this downregulation is achieved in the CMM is unknown. Here, we have studied the role of HISTONE DEACETYLASE 19 (HDA19) in Arabidopsis (Arabidopsis thaliana) during ovule and ReT differentiation based on the observation that the hda19-3 mutant displays a reduced ovule number and fails to differentiate the TT properly. Fluorescence-activated cell sorting coupled with RNA-sequencing revealed that in the CMM of hda19-3 mutants, genes promoting organ development are downregulated while meristematic markers, including STM, are upregulated. HDA19 was essential to downregulate STM in the CMM, thereby allowing ovule formation and TT differentiation. STM is ectopically expressed in hda19-3 at intermediate stages of pistil development, and its downregulation by RNA interference alleviated the hda19-3 phenotype. Chromatin immunoprecipitation assays indicated that STM is a direct target of HDA19 during pistil development and that the transcription factor SEEDSTICK is also required to regulate STM via histone acetylation. Thus, we identified factors required for the downregulation of STM in the CMM, which is necessary for organogenesis and tissue differentiation.S.Man., A.C., and L.C. were funded by MIUR (PRIN2017). A.G. and L.C. were funded by H2020 Excellent Science (SEXSEED ID: 690946). S.Man. was funded by an EMBO short-term fellowship (2015). S.Man., S.Mas., and C.M. were funded by FP7 People: Marie-Curie Actions International Research Staff Exchange Scheme (FRUITLOOK ID: 612640). S.C. was funded by Fundação para a Ciência e a Tecnologia (Project POCI-01-0145-FEDER-027839). U.G. was funded by the Swiss National Science Foundation (grant 31003A_179553).American Society of Plant PhysiologistsDepartamento de BiotecnologíaInstituto Universitario de Conservación y Mejora de la Agrodiversidad ValencianaEscuela Técnica Superior de Ingeniería Agronómica y del Medio NaturalEuropean CommissionSwiss National Science FoundationEuropean Molecular Biology OrganizationFundação para a Ciência e a Tecnologia, PortugalMinistero dell'Istruzione, dell'Università e della RicercaRepositorio Institucional de la Universitat Politècnica de València Riunet20242024-01-01journal articlehttp://purl.org/coar/resource_type/c_6501VoRhttp://purl.org/coar/version/c_970fb48d4fbd8a85info:eu-repo/semantics/articleapplication/pdfhttps://riunet.upv.es/handle/10251/234136reponame:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valénciainstname:Universitat Politècnica de València (UPV)InglésengEuropean Commission https://doi.org/10.13039/501100000780 FP7 612640 The physiology and genetics of fruit formation: from genes to networksEuropean Commission https://doi.org/10.13039/501100000780 H2020 690946 Sexual Plant Reproduction ¿ Seed formationFundação para a Ciência e a Tecnologia, Portugal https://doi.org/10.13039/501100001871 POCI-01-0145-FEDER-027839Swiss National Science Foundation https://doi.org/10.13039/501100001711 31003A_179553open accesshttp://purl.org/coar/access_right/c_abf2Reconocimiento (by)http://creativecommons.org/licenses/by/4.0/info:eu-repo/semantics/openAccessoai:dnet:riunet______::6b0a516ebfdf8c96bc387baf54e0ede32026-06-13T07:49:27Z
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