Basolateral sorting and transcytosis define the cu+-regulated translocation of ATP7B to the bile canaliculus

The Cu pump ATP7B plays an irreplaceable role in the elimination of excess Cu by the hepatocyte into the bile. The trafficking and site of action of ATP7B are subjects of controversy. One current proposal is that an increase in intracellular Cu results in the translocation of ATP7B to the lysosomes...

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Detalles Bibliográficos
Autores: Lalioti, Vasiliky S., Peiró-Pastor, Ramón, Tsuchiya, Yo, Muñoz, Ángeles, Villalba, Teresa, Sánchez, Carlos, Sandoval, Ignacio V., Pérez-Berlanga, Manuela
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2016
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/150712
Acceso en línea:http://hdl.handle.net/10261/150712
Access Level:acceso abierto
Palabra clave:Copper
Trafficking
TGN
Bile canaliculus
Transcytosis
ATP7B
Descripción
Sumario:The Cu pump ATP7B plays an irreplaceable role in the elimination of excess Cu by the hepatocyte into the bile. The trafficking and site of action of ATP7B are subjects of controversy. One current proposal is that an increase in intracellular Cu results in the translocation of ATP7B to the lysosomes and excretion of excess Cu through lysosomal-mediated exocytosis at the bile canaliculus. Here, we show that ATP7B is transported from the trans-Golgi network (TGN) to the bile canaliculus by basolateral sorting and endocytosis, and microtubule-mediated transcytosis through the subapical compartment. Trafficking ATP7B is not incorporated into lysosomes, and addition of Cu does not cause relocalization of lysosomes and the appearance of lysosome markers in the bile canaliculus. Our data reveal the pathway of the Cu-mediated transport of ATP7B from the TGN to the bile canaliculus and indicates that the bile canaliculus is the primary site of ATP7B action in the elimination of excess Cu.