Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast

The three-dimensional (3D) organization of chromosomes can influence transcription. However, the frequency and magnitude of these effects remain debated. To determine how changes in chromosome positioning affect transcription across thousands of genes with minimal perturbation, we characterized nucl...

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Autores: Di Stefano, Marco, Di Giovanni, Francesca, 1985-, Pozharskaia, Vasilia, Gomar-Alba, Mercè, Baù, Davide, Carey, Lucas, 1980-, Marti-Renom, Marc A., Mendoza, Manuel (Mendoza Palomares)
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2020
País:España
Institución:Universitat Pompeu Fabra
Repositorio:Repositorio Digital de la UPF
OAI Identifier:oai:repositori.upf.edu:10230/43995
Acceso en línea:http://hdl.handle.net/10230/43995
http://dx.doi.org/10.1534/genetics.119.302978
Access Level:acceso abierto
Palabra clave:Cromosomes
Genomes
Expressió gènica
Llevats
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network_name_str España
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dc.title.none.fl_str_mv Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
title Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
spellingShingle Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
Di Stefano, Marco
Cromosomes
Genomes
Expressió gènica
Llevats
title_short Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
title_full Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
title_fullStr Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
title_full_unstemmed Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
title_sort Impact of chromosome fusions on 3D genome organization and gene expression in budding yeast
dc.creator.none.fl_str_mv Di Stefano, Marco
Di Giovanni, Francesca, 1985-
Pozharskaia, Vasilia
Gomar-Alba, Mercè
Baù, Davide
Carey, Lucas, 1980-
Marti-Renom, Marc A.
Mendoza, Manuel (Mendoza Palomares)
author Di Stefano, Marco
author_facet Di Stefano, Marco
Di Giovanni, Francesca, 1985-
Pozharskaia, Vasilia
Gomar-Alba, Mercè
Baù, Davide
Carey, Lucas, 1980-
Marti-Renom, Marc A.
Mendoza, Manuel (Mendoza Palomares)
author_role author
author2 Di Giovanni, Francesca, 1985-
Pozharskaia, Vasilia
Gomar-Alba, Mercè
Baù, Davide
Carey, Lucas, 1980-
Marti-Renom, Marc A.
Mendoza, Manuel (Mendoza Palomares)
author2_role author
author
author
author
author
author
author
dc.subject.none.fl_str_mv Cromosomes
Genomes
Expressió gènica
Llevats
topic Cromosomes
Genomes
Expressió gènica
Llevats
description The three-dimensional (3D) organization of chromosomes can influence transcription. However, the frequency and magnitude of these effects remain debated. To determine how changes in chromosome positioning affect transcription across thousands of genes with minimal perturbation, we characterized nuclear organization and global gene expression in budding yeast containing chromosome fusions. We used computational modeling and single-cell imaging to determine chromosome positions, and integrated these data with genome-wide transcriptional profiles from RNA sequencing. We find that chromosome fusions dramatically alter 3D nuclear organization without leading to strong genome-wide changes in transcription. However, we observe a mild but significant and reproducible increase in the expression of genes displaced away from the periphery. The increase in transcription is inversely proportional to the propensity of a given locus to be at the nuclear periphery; for example, a 10% decrease in the propensity of a gene to reside at the nuclear envelope is accompanied by a 10% increase in gene expression. Modeling suggests that this is due to both deletion of telomeres and to displacement of genes relative to the nuclear periphery. These data suggest that basal transcriptional activity is sensitive to radial changes in gene position, and provide insight into the functional relevance of budding yeast chromosome-level 3D organization in gene expression.
publishDate 2020
dc.date.none.fl_str_mv 2020
2020
2020
dc.type.none.fl_str_mv info:eu-repo/semantics/article
info:eu-repo/semantics/publishedVersion
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv http://hdl.handle.net/10230/43995
http://dx.doi.org/10.1534/genetics.119.302978
url http://hdl.handle.net/10230/43995
http://dx.doi.org/10.1534/genetics.119.302978
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
dc.relation.none.fl_str_mv Genetics. 2020 Mar; 214(3) :651-67
info:eu-repo/grantAgreement/EC/H2020/676556
info:eu-repo/grantAgreement/ES/3PN/BFU2012-37162
info:eu-repo/grantAgreement/ES/1PE/BFU2015-68351-P
info:eu-repo/grantAgreement/ES/2PE/BFU2017-85926-P
dc.rights.none.fl_str_mv http://creativecommons.org/licenses/by/4.0/
info:eu-repo/semantics/openAccess
rights_invalid_str_mv http://creativecommons.org/licenses/by/4.0/
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
application/pdf
dc.publisher.none.fl_str_mv Genetics Society of America
publisher.none.fl_str_mv Genetics Society of America
dc.source.none.fl_str_mv reponame:Repositorio Digital de la UPF
instname:Universitat Pompeu Fabra
instname_str Universitat Pompeu Fabra
reponame_str Repositorio Digital de la UPF
collection Repositorio Digital de la UPF
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spelling Impact of chromosome fusions on 3D genome organization and gene expression in budding yeastDi Stefano, MarcoDi Giovanni, Francesca, 1985-Pozharskaia, VasiliaGomar-Alba, MercèBaù, DavideCarey, Lucas, 1980-Marti-Renom, Marc A.Mendoza, Manuel (Mendoza Palomares)CromosomesGenomesExpressió gènicaLlevatsThe three-dimensional (3D) organization of chromosomes can influence transcription. However, the frequency and magnitude of these effects remain debated. To determine how changes in chromosome positioning affect transcription across thousands of genes with minimal perturbation, we characterized nuclear organization and global gene expression in budding yeast containing chromosome fusions. We used computational modeling and single-cell imaging to determine chromosome positions, and integrated these data with genome-wide transcriptional profiles from RNA sequencing. We find that chromosome fusions dramatically alter 3D nuclear organization without leading to strong genome-wide changes in transcription. However, we observe a mild but significant and reproducible increase in the expression of genes displaced away from the periphery. The increase in transcription is inversely proportional to the propensity of a given locus to be at the nuclear periphery; for example, a 10% decrease in the propensity of a gene to reside at the nuclear envelope is accompanied by a 10% increase in gene expression. Modeling suggests that this is due to both deletion of telomeres and to displacement of genes relative to the nuclear periphery. These data suggest that basal transcriptional activity is sensitive to radial changes in gene position, and provide insight into the functional relevance of budding yeast chromosome-level 3D organization in gene expression.This project was supported by European Research Council Starting grant 2010-St-20091118 and Spanish Ministry of Economy and Competitiveness grant BFU2012-37162 to M.M., and grant ANR-10-LABX-0030-INRT, which is a French State fund managed by the Agence Nationale de la Recherche under the frame Programme Investissements d’Avenir ANR-10-IDEX-0002-02 to the Institut de Génétique et de Biologie Moléculaire et Cellulaire. L.B.C. was supported by the Agència de Gestió d’Ajuts Universitaris i de Recerca (grant 2014 SGR 0974), and the Spanish Ministerio de Economía y Competitividad and the European Regional Development Fund (FEDER) (grant BFU2015-68351-P). M.A.M.-R. was supported by the European Union’s Horizon 2020 research and innovation programme (grant agreement 676556) and the Spanish Ministry of Economy and Competitiveness (grant BFU2017-85926-P). We also acknowledge the support of the Spanish Ministry of Science and Innovation to the EMBL partnership, the ‘Centro de Excelencia Severo Ochoa 2013-2017’, SEV-2012-0208, the CERCA Programme/Generalitat de Catalunya, Spanish Ministry of Science and Innovation through the Instituto de Salud Carlos III, the Generalitat de Catalunya through Departament de Salut and Departament d’Empresa i Coneixement and the Co-financing by Spanish Ministry of Science and Innovation with funds from the European Regional Development Fund (ERDF) corresponding to the 2014-2020 Smart Growth Operating Program to the CRG. M.G.-A. is a recipient of a Postdoctoral Fellowship (APOSTD/2017/094) from the Generalitat ValencianaGenetics Society of America202020202020info:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionapplication/pdfapplication/pdfhttp://hdl.handle.net/10230/43995http://dx.doi.org/10.1534/genetics.119.302978reponame:Repositorio Digital de la UPFinstname:Universitat Pompeu FabraInglésGenetics. 2020 Mar; 214(3) :651-67info:eu-repo/grantAgreement/EC/H2020/676556info:eu-repo/grantAgreement/ES/3PN/BFU2012-37162info:eu-repo/grantAgreement/ES/1PE/BFU2015-68351-Pinfo:eu-repo/grantAgreement/ES/2PE/BFU2017-85926-P© 2020 Marco Di Stefano et al. This is an open-access article distributed under the terms of the Creative Commons Attribution 4.0 International License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly citedhttp://creativecommons.org/licenses/by/4.0/info:eu-repo/semantics/openAccessoai:repositori.upf.edu:10230/439952026-06-12T07:21:37Z
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