Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs

1. Competitive and facilitative interactions shape plant communities. Whereas a number of studies have addressed competition and direct facilitation among plants in dry ecosystems, indirect facilitation has received little attention. 2. We investigated the relative importance of direct and indirect...

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Autores: Cuesta Poveda, Bárbara, Villar Salvador, Pedro|||0000-0001-9338-4530, Rey Benayas, José María|||0000-0002-2099-8512, Puértolas Simón, Jaime, Michalet, Richard
Formato: artículo
Fecha de publicación:2010
País:España
Recursos:Universidad de Alcalá (UAH)
Repositorio:e_Buah Biblioteca Digital Universidad de Alcalá
Idioma:inglés
OAI Identifier:oai:ebuah.uah.es:10017/21314
Acesso em linha:http://hdl.handle.net/10017/21314
https://dx.doi.org/10.1111/j.1365-2745.2010.01655.x
Access Level:acceso abierto
Palavra-chave:Chlorophyll fluorescence
Herb competition
Quercus ilex
Retama sphaerocarpa
Seedling size
Water potential
Ciencia
Medio Ambiente
Science
Environmental science
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spelling Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbsCuesta Poveda, BárbaraVillar Salvador, Pedro|||0000-0001-9338-4530Rey Benayas, José María|||0000-0002-2099-8512Puértolas Simón, JaimeMichalet, RichardChlorophyll fluorescenceHerb competitionQuercus ilexRetama sphaerocarpaSeedling sizeWater potentialCienciaMedio AmbienteScienceEnvironmental science1. Competitive and facilitative interactions shape plant communities. Whereas a number of studies have addressed competition and direct facilitation among plants in dry ecosystems, indirect facilitation has received little attention. 2. We investigated the relative importance of direct and indirect facilitation by the nurse plant Retama sphaerocarpa on late-successional Quercus ilex seedlings mediated by herb suppression in a Mediterranean shrubland in 2006 and 2007. We also studied whether facilitation outcome depended on the size of the facilitated seedlings. 3. A field experiment was carried out to test the effect of (i) position of Q. ilex seedling with respect to shrub canopy (under shrubs or in gaps), (ii) herb competition (presence or absence), and (iii) seedling size. 2006 was an average rainfall year while 2007 had a much more humid spring and a dryer summer than 2006. 4. In both years, nurse shrubs reduced seedling mortality whereas herbs increased it. In the average rainfall year, seedling mortality under shrubs was unaffected by herbs whereas in gaps it was signifi- cantly higher in presence of herbs. This showed that the nurse shrub indirectly facilitated the seedlings by reducing the competitive capacity of herbs. Conversely, facilitation was predominately direct during the humid spring and dry summer year since herbs hindered seedling survival similarly under the nurse shrub and in gaps. The nurse shrub directly facilitated the seedlings by reducing seedling photoinhibition and water stress. 5. Improvement of environmental conditions by Retama benefited smaller seedlings but not larger seedlings since the nurse shrub reduced mortality of smaller seedlings relative to that in gaps, but this effect was not observed for larger seedlings. This indicates that individuals within a seedling population may not have the same response to facilitation. 6. Synthesis. Both indirect and direct facilitation are important mechanisms for Q. ilex regeneration in Retama shrubland and their importance seems to vary with climatic conditions. Indirect facilitation by release of herb competition under nurse shrubs is important in years of dry springs when competition between nurse shrubs and herbs is high, whereas direct facilitation mediated by microclimate amelioration increases with summer aridityMinisterio de Educación y CienciaMinisterio de Ciencia y TecnologíaComunidad de MadridJohn Wiley & Sons20102010-01-01journal articlehttp://purl.org/coar/resource_type/c_6501NAhttp://purl.org/coar/version/c_be7fb7dd8ff6fe43info:eu-repo/semantics/articleapplication/pdfhttp://hdl.handle.net/10017/21314https://dx.doi.org/10.1111/j.1365-2745.2010.01655.xreponame:e_Buah Biblioteca Digital Universidad de Alcaláinstname:Universidad de Alcalá (UAH)InglésengMinisterio de Educación y Ciencia Not available CGL2007-60533 EFECTOS DE LA REVEGETACION (PASIVA Y ACTIVA) EN LA DINAMICA Y DIVERSIDAD DE ESPECIES LEÑOSAS Y AVESMCICYT Not available AGL2006-12609-C02-01%2FFOR Optimización de la nutrición de la encina en vivero: bases fisiológicas, técnicas de fertilización e implicaciones post-transplanteComunidad de Madrid http://dx.doi.org/10.13039/100012818 Not available S0505%2FAMB%2F0355 Programa de actividades de I+D de la Comunidad de Madrid sobre restauración del medio naturalopen accesshttp://purl.org/coar/access_right/c_abf2Atribución-NoComercial-SinDerivadas 3.0 Españahttp://creativecommons.org/licenses/by-nc-nd/3.0/es/info:eu-repo/semantics/openAccessoai:ebuah.uah.es:10017/213142026-06-18T11:13:07Z
dc.title.none.fl_str_mv Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
title Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
spellingShingle Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
Cuesta Poveda, Bárbara
Chlorophyll fluorescence
Herb competition
Quercus ilex
Retama sphaerocarpa
Seedling size
Water potential
Ciencia
Medio Ambiente
Science
Environmental science
title_short Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
title_full Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
title_fullStr Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
title_full_unstemmed Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
title_sort Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs
dc.creator.none.fl_str_mv Cuesta Poveda, Bárbara
Villar Salvador, Pedro|||0000-0001-9338-4530
Rey Benayas, José María|||0000-0002-2099-8512
Puértolas Simón, Jaime
Michalet, Richard
author Cuesta Poveda, Bárbara
author_facet Cuesta Poveda, Bárbara
Villar Salvador, Pedro|||0000-0001-9338-4530
Rey Benayas, José María|||0000-0002-2099-8512
Puértolas Simón, Jaime
Michalet, Richard
author_role author
author2 Villar Salvador, Pedro|||0000-0001-9338-4530
Rey Benayas, José María|||0000-0002-2099-8512
Puértolas Simón, Jaime
Michalet, Richard
author2_role author
author
author
author
dc.subject.none.fl_str_mv Chlorophyll fluorescence
Herb competition
Quercus ilex
Retama sphaerocarpa
Seedling size
Water potential
Ciencia
Medio Ambiente
Science
Environmental science
topic Chlorophyll fluorescence
Herb competition
Quercus ilex
Retama sphaerocarpa
Seedling size
Water potential
Ciencia
Medio Ambiente
Science
Environmental science
description 1. Competitive and facilitative interactions shape plant communities. Whereas a number of studies have addressed competition and direct facilitation among plants in dry ecosystems, indirect facilitation has received little attention. 2. We investigated the relative importance of direct and indirect facilitation by the nurse plant Retama sphaerocarpa on late-successional Quercus ilex seedlings mediated by herb suppression in a Mediterranean shrubland in 2006 and 2007. We also studied whether facilitation outcome depended on the size of the facilitated seedlings. 3. A field experiment was carried out to test the effect of (i) position of Q. ilex seedling with respect to shrub canopy (under shrubs or in gaps), (ii) herb competition (presence or absence), and (iii) seedling size. 2006 was an average rainfall year while 2007 had a much more humid spring and a dryer summer than 2006. 4. In both years, nurse shrubs reduced seedling mortality whereas herbs increased it. In the average rainfall year, seedling mortality under shrubs was unaffected by herbs whereas in gaps it was signifi- cantly higher in presence of herbs. This showed that the nurse shrub indirectly facilitated the seedlings by reducing the competitive capacity of herbs. Conversely, facilitation was predominately direct during the humid spring and dry summer year since herbs hindered seedling survival similarly under the nurse shrub and in gaps. The nurse shrub directly facilitated the seedlings by reducing seedling photoinhibition and water stress. 5. Improvement of environmental conditions by Retama benefited smaller seedlings but not larger seedlings since the nurse shrub reduced mortality of smaller seedlings relative to that in gaps, but this effect was not observed for larger seedlings. This indicates that individuals within a seedling population may not have the same response to facilitation. 6. Synthesis. Both indirect and direct facilitation are important mechanisms for Q. ilex regeneration in Retama shrubland and their importance seems to vary with climatic conditions. Indirect facilitation by release of herb competition under nurse shrubs is important in years of dry springs when competition between nurse shrubs and herbs is high, whereas direct facilitation mediated by microclimate amelioration increases with summer aridity
publishDate 2010
dc.date.none.fl_str_mv 2010
2010-01-01
dc.type.none.fl_str_mv journal article
http://purl.org/coar/resource_type/c_6501
NA
http://purl.org/coar/version/c_be7fb7dd8ff6fe43
dc.type.openaire.fl_str_mv info:eu-repo/semantics/article
format article
dc.identifier.none.fl_str_mv http://hdl.handle.net/10017/21314
https://dx.doi.org/10.1111/j.1365-2745.2010.01655.x
url http://hdl.handle.net/10017/21314
https://dx.doi.org/10.1111/j.1365-2745.2010.01655.x
dc.language.none.fl_str_mv Inglés
eng
language_invalid_str_mv Inglés
language eng
dc.relation.none.fl_str_mv Ministerio de Educación y Ciencia Not available CGL2007-60533 EFECTOS DE LA REVEGETACION (PASIVA Y ACTIVA) EN LA DINAMICA Y DIVERSIDAD DE ESPECIES LEÑOSAS Y AVES
MCICYT Not available AGL2006-12609-C02-01%2FFOR Optimización de la nutrición de la encina en vivero: bases fisiológicas, técnicas de fertilización e implicaciones post-transplante
Comunidad de Madrid http://dx.doi.org/10.13039/100012818 Not available S0505%2FAMB%2F0355 Programa de actividades de I+D de la Comunidad de Madrid sobre restauración del medio natural
dc.rights.none.fl_str_mv open access
http://purl.org/coar/access_right/c_abf2
Atribución-NoComercial-SinDerivadas 3.0 España
http://creativecommons.org/licenses/by-nc-nd/3.0/es/
dc.rights.openaire.fl_str_mv info:eu-repo/semantics/openAccess
rights_invalid_str_mv open access
http://purl.org/coar/access_right/c_abf2
Atribución-NoComercial-SinDerivadas 3.0 España
http://creativecommons.org/licenses/by-nc-nd/3.0/es/
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
dc.publisher.none.fl_str_mv John Wiley & Sons
publisher.none.fl_str_mv John Wiley & Sons
dc.source.none.fl_str_mv reponame:e_Buah Biblioteca Digital Universidad de Alcalá
instname:Universidad de Alcalá (UAH)
instname_str Universidad de Alcalá (UAH)
reponame_str e_Buah Biblioteca Digital Universidad de Alcalá
collection e_Buah Biblioteca Digital Universidad de Alcalá
repository.name.fl_str_mv
repository.mail.fl_str_mv
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