GIGANTEA adjusts the response to shade at dusk by directly impinging on PHYTOCHROME INTERACTING FACTOR 7 function

[EN] For plants adapted to bright light, a decrease in the amount of light received can be detrimental to their growth and survival. Consequently, in response to shade from surrounding vegetation, they initiate a suite of molecular and morphological changes known as the shade avoidance response thro...

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Detalles Bibliográficos
Autores: Martínez-Vasallo, Carlos, Cole, Benjamin, Pérez-Castillo, Jaime, Chory, Joanne, Kay, Steve A., Ortiz-Ramirez, Clara Inés, Gallego Bartolomé, Javier, Nohales, Maria A.
Tipo de recurso: artículo
Fecha de publicación:2024
País:España
Institución:Universitat Politècnica de València (UPV)
Repositorio:RiuNet. Repositorio Institucional de la Universitat Politécnica de Valéncia
Idioma:inglés
OAI Identifier:oai:riunet.upv.es:10251/214764
Acceso en línea:https://riunet.upv.es/handle/10251/214764
Access Level:acceso abierto
Palabra clave:Shade avoidance
Circadian gating
GIGANTEA
PIF7
Tissue specificity
Descripción
Sumario:[EN] For plants adapted to bright light, a decrease in the amount of light received can be detrimental to their growth and survival. Consequently, in response to shade from surrounding vegetation, they initiate a suite of molecular and morphological changes known as the shade avoidance response through which stems and petioles elongate in search for light. Under sunlight-night cycles, the plant's responsiveness to shade varies across the day, being maximal at dusk time. While a role for the circadian clock in this regulation has long been proposed, mechanistic understanding of how it is achieved is incomplete. Here, we show that the clock component GIGANTEA (GI) directly interacts with the key player in the response to shade. GI represses PIF7 transcriptional activity and the expression of its target genes in response to shade, thereby fine- tuning the magnitude of the response to limiting light conditions. We find that under light/dark cycles, this dusk. Importantly, we also show that this circuit primarily operates in epidermal cells, highlighting the relevance of tissue- specific clock- output connections for the regulation of plant development in resonance with the environment.