Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains

[EN] The binding and early stages of activity of a phospholipase C/sphingomyelinase from Pseudomonas aeruginosa on giant unilamellar vesicles (GUV) have been monitored using fl uorescence confocal microscopy. Both the lipids and the enzyme were labeled with specifi c fl uorescent markers. GUV consis...

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Autores: Ibarguren, Maitane, López Jiménez, David, Montes Burgos, Lidia Ruth, Sot Sanz, Jesús, Vasil, Michael L., Vasil, Adriana I., Goñi Urcelay, Félix María, Alonso Izquierdo, Alicia
Tipo de recurso: artículo
Fecha de publicación:2011
País:España
Institución:Universidad del País Vasco
Repositorio:Addi. Archivo Digital para la Docencia y la Investigación
OAI Identifier:oai:addi.ehu.eus:10810/64693
Acceso en línea:http://hdl.handle.net/10810/64693
Access Level:acceso abierto
Palabra clave:ceramides
cholesterol
diacylglycerol
fluorescence microscopy
lipid rafts
membranes/physical chemistry
sphingolipids
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oai_identifier_str oai:addi.ehu.eus:10810/64693
network_acronym_str ES
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repository_id_str
spelling Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domainsIbarguren, MaitaneLópez Jiménez, DavidMontes Burgos, Lidia RuthSot Sanz, JesúsVasil, Michael L.Vasil, Adriana I.Goñi Urcelay, Félix MaríaAlonso Izquierdo, Aliciaceramidescholesteroldiacylglycerolfluorescence microscopylipid raftsmembranes/physical chemistrysphingolipids[EN] The binding and early stages of activity of a phospholipase C/sphingomyelinase from Pseudomonas aeruginosa on giant unilamellar vesicles (GUV) have been monitored using fl uorescence confocal microscopy. Both the lipids and the enzyme were labeled with specifi c fl uorescent markers. GUV consisted of a mixture of phosphatidylcholine, sphingomyelin,phosphatidylethanolamine, and cholesterol in equimolar ratios, to which 5–10 mol% of the enzyme endproduct ceramide and/or diacylglycerol were occasionally added. Morphological examination of the GUV in the presence of enzyme reveals that, although the enzyme diffuses rapidly throughout the observation chamber, detectable enzyme binding appears to be a slow, random process, with new bound-enzyme-containing vesicles appearing for several minutes. Enzyme binding to the vesicles appears to be a cooperative process. After the initial cluster of bound enzyme is detected, further binding and catalytic activity follow rapidly. After the activity has started, the enzyme is not released by repeated washing, suggesting a “scooting” mechanism for the hydrolytic activity. The enzyme preferentially binds the more disordered domains, and, in most cases, the catalytic activity causes the disordering of the other domains. Simultaneously, peanut- or fi gure-eight-shaped vesicles containing two separate lipid domains become spherical. At a further stage of lipid hydrolysis, lipid aggregates are formed and vesicles disintegrate.Spanish Ministerio de Ciencia e Innovación Grants BFU 2008-01637/BMC and BFU 2007-62062 Basque Government Grant IT 461-07 ETORTEK 07/26 National Institutes of Health Grant HL-062608 Basque Government.Elsevier202420242011info:eu-repo/semantics/articleapplication/pdfhttp://hdl.handle.net/10810/64693reponame:Addi. Archivo Digital para la Docencia y la Investigacióninstname:Universidad del País VascoIngléshttps://www.sciencedirect.com/science/article/pii/S0022227520408983?via%3Dihubinfo:eu-repo/semantics/openAccesshttp://creativecommons.org/licenses/by/4.0/© 2011 by the American Society for Biochemistry and Molecular Biology, Inc. This is an Open Access article under the CC BY licenseoai:addi.ehu.eus:10810/646932026-06-18T09:23:17Z
dc.title.none.fl_str_mv Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
title Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
spellingShingle Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
Ibarguren, Maitane
ceramides
cholesterol
diacylglycerol
fluorescence microscopy
lipid rafts
membranes/physical chemistry
sphingolipids
title_short Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
title_full Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
title_fullStr Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
title_full_unstemmed Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
title_sort Imaging the early stages of phospholipase C/sphingomyelinase activity on vesicles containing coexisting order-disorder and gel-fluid domains
dc.creator.none.fl_str_mv Ibarguren, Maitane
López Jiménez, David
Montes Burgos, Lidia Ruth
Sot Sanz, Jesús
Vasil, Michael L.
Vasil, Adriana I.
Goñi Urcelay, Félix María
Alonso Izquierdo, Alicia
author Ibarguren, Maitane
author_facet Ibarguren, Maitane
López Jiménez, David
Montes Burgos, Lidia Ruth
Sot Sanz, Jesús
Vasil, Michael L.
Vasil, Adriana I.
Goñi Urcelay, Félix María
Alonso Izquierdo, Alicia
author_role author
author2 López Jiménez, David
Montes Burgos, Lidia Ruth
Sot Sanz, Jesús
Vasil, Michael L.
Vasil, Adriana I.
Goñi Urcelay, Félix María
Alonso Izquierdo, Alicia
author2_role author
author
author
author
author
author
author
dc.subject.none.fl_str_mv ceramides
cholesterol
diacylglycerol
fluorescence microscopy
lipid rafts
membranes/physical chemistry
sphingolipids
topic ceramides
cholesterol
diacylglycerol
fluorescence microscopy
lipid rafts
membranes/physical chemistry
sphingolipids
description [EN] The binding and early stages of activity of a phospholipase C/sphingomyelinase from Pseudomonas aeruginosa on giant unilamellar vesicles (GUV) have been monitored using fl uorescence confocal microscopy. Both the lipids and the enzyme were labeled with specifi c fl uorescent markers. GUV consisted of a mixture of phosphatidylcholine, sphingomyelin,phosphatidylethanolamine, and cholesterol in equimolar ratios, to which 5–10 mol% of the enzyme endproduct ceramide and/or diacylglycerol were occasionally added. Morphological examination of the GUV in the presence of enzyme reveals that, although the enzyme diffuses rapidly throughout the observation chamber, detectable enzyme binding appears to be a slow, random process, with new bound-enzyme-containing vesicles appearing for several minutes. Enzyme binding to the vesicles appears to be a cooperative process. After the initial cluster of bound enzyme is detected, further binding and catalytic activity follow rapidly. After the activity has started, the enzyme is not released by repeated washing, suggesting a “scooting” mechanism for the hydrolytic activity. The enzyme preferentially binds the more disordered domains, and, in most cases, the catalytic activity causes the disordering of the other domains. Simultaneously, peanut- or fi gure-eight-shaped vesicles containing two separate lipid domains become spherical. At a further stage of lipid hydrolysis, lipid aggregates are formed and vesicles disintegrate.
publishDate 2011
dc.date.none.fl_str_mv 2011
2024
2024
dc.type.none.fl_str_mv info:eu-repo/semantics/article
format article
dc.identifier.none.fl_str_mv http://hdl.handle.net/10810/64693
url http://hdl.handle.net/10810/64693
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
dc.relation.none.fl_str_mv https://www.sciencedirect.com/science/article/pii/S0022227520408983?via%3Dihub
dc.rights.none.fl_str_mv info:eu-repo/semantics/openAccess
http://creativecommons.org/licenses/by/4.0/
eu_rights_str_mv openAccess
rights_invalid_str_mv http://creativecommons.org/licenses/by/4.0/
dc.format.none.fl_str_mv application/pdf
dc.publisher.none.fl_str_mv Elsevier
publisher.none.fl_str_mv Elsevier
dc.source.none.fl_str_mv reponame:Addi. Archivo Digital para la Docencia y la Investigación
instname:Universidad del País Vasco
instname_str Universidad del País Vasco
reponame_str Addi. Archivo Digital para la Docencia y la Investigación
collection Addi. Archivo Digital para la Docencia y la Investigación
repository.name.fl_str_mv
repository.mail.fl_str_mv
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