SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans

Dectin-1 (Clec7a) is a paradigmatic C-type lectin receptor that binds Syk through a hemITAM motif and couples sensing of pathogens such as fungi to induction of innate responses. Dectin-1 engagement triggers a plethora of activating events, but little is known about the modulation of such pathways....

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Detalles Bibliográficos
Autores: Blanco-Menendez, Noelia, del Fresno, Carlos, Fernandes, Sandra, Calvo, Enrique, Conde-Garrosa, Ruth, Kerr, William G, Sancho, David
Tipo de recurso: artículo
Fecha de publicación:2015
País:España
Institución:Instituto de Salud Carlos III (ISCIII)
Repositorio:Repisalud
Idioma:inglés
OAI Identifier:oai:repisalud.isciii.es:20.500.12105/9706
Acceso en línea:http://hdl.handle.net/20.500.12105/9706
Access Level:acceso abierto
Palabra clave:Amino Acid Motifs
Amino Acid Sequence
Animals
Blotting, Western
Bone Marrow Cells
Candida albicans
Dendritic Cells
Granulocyte-Macrophage Colony-Stimulating Factor
Host-Pathogen Interactions
Inositol Polyphosphate 5-Phosphatases
Lectins, C-Type
Macrophages
Mice, Knockout
Microscopy, Confocal
Molecular Sequence Data
NF-kappa B
NFATC Transcription Factors
Phagocytosis
Phagosomes
Phosphatidylinositol-3,4,5-Trisphosphate 5-Phosphatases
Phosphoric Monoester Hydrolases
Protein Binding
Reactive Oxygen Species
Signal Transduction
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spelling SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicansBlanco-Menendez, Noeliadel Fresno, CarlosFernandes, SandraCalvo, EnriqueConde-Garrosa, RuthKerr, William GSancho, DavidAmino Acid MotifsAmino Acid SequenceAnimalsBlotting, WesternBone Marrow CellsCandida albicansDendritic CellsGranulocyte-Macrophage Colony-Stimulating FactorHost-Pathogen InteractionsInositol Polyphosphate 5-PhosphatasesLectins, C-TypeMacrophagesMice, KnockoutMicroscopy, ConfocalMolecular Sequence DataNF-kappa BNFATC Transcription FactorsPhagocytosisPhagosomesPhosphatidylinositol-3,4,5-Trisphosphate 5-PhosphatasesPhosphoric Monoester HydrolasesProtein BindingReactive Oxygen SpeciesSignal TransductionDectin-1 (Clec7a) is a paradigmatic C-type lectin receptor that binds Syk through a hemITAM motif and couples sensing of pathogens such as fungi to induction of innate responses. Dectin-1 engagement triggers a plethora of activating events, but little is known about the modulation of such pathways. Trying to define a more precise picture of early Dectin-1 signaling, we explored the interactome of the intracellular tail of the receptor in mouse dendritic cells. We found unexpected binding of SHIP-1 phosphatase to the phosphorylated hemITAM. SHIP-1 colocalized with Dectin-1 during phagocytosis of zymosan in a hemITAM-dependent fashion. Moreover, endogenous SHIP-1 relocated to live or heat-killed Candida albicans-containing phagosomes in a Dectin-1-dependent manner in GM-CSF-derived bone marrow cells (GM-BM). However, SHIP-1 absence in GM-BM did not affect activation of MAPK or production of cytokines and readouts dependent on NF-κB and NFAT. Notably, ROS production was enhanced in SHIP-1-deficient GM-BM treated with heat-killed C. albicans, live C. albicans, or the specific Dectin-1 agonists curdlan or whole glucan particles. This increased oxidative burst was dependent on Dectin-1, Syk, PI3K, phosphoinositide-dependent protein kinase 1, and NADPH oxidase. GM-BM from CD11c∆SHIP-1 mice also showed increased killing activity against live C. albicans that was dependent on Dectin-1, Syk, and NADPH oxidase. These results illustrate the complexity of myeloid C-type lectin receptor signaling, and how an activating hemITAM can also couple to intracellular inositol phosphatases to modulate selected functional responses and tightly regulate processes such as ROS production that could be deleterious to the host.American Association of Immunologists (AAI)Ministerio de Economía y Competitividad (España)Unión Europea. Comisión Europea. European Research Council (ERC)Fundación ProCNICNational Institutes of Health (Estados Unidos)20202020-04-2320152015-11-0120152015-11-01journal articlehttp://purl.org/coar/resource_type/c_6501AMhttp://purl.org/coar/version/c_ab4af688f83e57aainfo:eu-repo/semantics/articleapplication/pdfhttp://hdl.handle.net/20.500.12105/9706reponame:Repisaludinstname:Instituto de Salud Carlos III (ISCIII)Inglésengopen accesshttp://purl.org/coar/access_right/c_abf2Attribution-NonCommercial-NoDerivatives 4.0 Internacionalhttp://creativecommons.org/licenses/by-nc-nd/4.0/info:eu-repo/semantics/openAccessoai:repisalud.isciii.es:20.500.12105/97062026-06-12T12:43:37Z
dc.title.none.fl_str_mv SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
title SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
spellingShingle SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
Blanco-Menendez, Noelia
Amino Acid Motifs
Amino Acid Sequence
Animals
Blotting, Western
Bone Marrow Cells
Candida albicans
Dendritic Cells
Granulocyte-Macrophage Colony-Stimulating Factor
Host-Pathogen Interactions
Inositol Polyphosphate 5-Phosphatases
Lectins, C-Type
Macrophages
Mice, Knockout
Microscopy, Confocal
Molecular Sequence Data
NF-kappa B
NFATC Transcription Factors
Phagocytosis
Phagosomes
Phosphatidylinositol-3,4,5-Trisphosphate 5-Phosphatases
Phosphoric Monoester Hydrolases
Protein Binding
Reactive Oxygen Species
Signal Transduction
title_short SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
title_full SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
title_fullStr SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
title_full_unstemmed SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
title_sort SHIP-1 Couples to the Dectin-1 hemITAM and Selectively Modulates Reactive Oxygen Species Production in Dendritic Cells in Response to Candida albicans
dc.creator.none.fl_str_mv Blanco-Menendez, Noelia
del Fresno, Carlos
Fernandes, Sandra
Calvo, Enrique
Conde-Garrosa, Ruth
Kerr, William G
Sancho, David
author Blanco-Menendez, Noelia
author_facet Blanco-Menendez, Noelia
del Fresno, Carlos
Fernandes, Sandra
Calvo, Enrique
Conde-Garrosa, Ruth
Kerr, William G
Sancho, David
author_role author
author2 del Fresno, Carlos
Fernandes, Sandra
Calvo, Enrique
Conde-Garrosa, Ruth
Kerr, William G
Sancho, David
author2_role author
author
author
author
author
author
dc.contributor.none.fl_str_mv Ministerio de Economía y Competitividad (España)
Unión Europea. Comisión Europea. European Research Council (ERC)
Fundación ProCNIC
National Institutes of Health (Estados Unidos)

dc.subject.none.fl_str_mv Amino Acid Motifs
Amino Acid Sequence
Animals
Blotting, Western
Bone Marrow Cells
Candida albicans
Dendritic Cells
Granulocyte-Macrophage Colony-Stimulating Factor
Host-Pathogen Interactions
Inositol Polyphosphate 5-Phosphatases
Lectins, C-Type
Macrophages
Mice, Knockout
Microscopy, Confocal
Molecular Sequence Data
NF-kappa B
NFATC Transcription Factors
Phagocytosis
Phagosomes
Phosphatidylinositol-3,4,5-Trisphosphate 5-Phosphatases
Phosphoric Monoester Hydrolases
Protein Binding
Reactive Oxygen Species
Signal Transduction
topic Amino Acid Motifs
Amino Acid Sequence
Animals
Blotting, Western
Bone Marrow Cells
Candida albicans
Dendritic Cells
Granulocyte-Macrophage Colony-Stimulating Factor
Host-Pathogen Interactions
Inositol Polyphosphate 5-Phosphatases
Lectins, C-Type
Macrophages
Mice, Knockout
Microscopy, Confocal
Molecular Sequence Data
NF-kappa B
NFATC Transcription Factors
Phagocytosis
Phagosomes
Phosphatidylinositol-3,4,5-Trisphosphate 5-Phosphatases
Phosphoric Monoester Hydrolases
Protein Binding
Reactive Oxygen Species
Signal Transduction
description Dectin-1 (Clec7a) is a paradigmatic C-type lectin receptor that binds Syk through a hemITAM motif and couples sensing of pathogens such as fungi to induction of innate responses. Dectin-1 engagement triggers a plethora of activating events, but little is known about the modulation of such pathways. Trying to define a more precise picture of early Dectin-1 signaling, we explored the interactome of the intracellular tail of the receptor in mouse dendritic cells. We found unexpected binding of SHIP-1 phosphatase to the phosphorylated hemITAM. SHIP-1 colocalized with Dectin-1 during phagocytosis of zymosan in a hemITAM-dependent fashion. Moreover, endogenous SHIP-1 relocated to live or heat-killed Candida albicans-containing phagosomes in a Dectin-1-dependent manner in GM-CSF-derived bone marrow cells (GM-BM). However, SHIP-1 absence in GM-BM did not affect activation of MAPK or production of cytokines and readouts dependent on NF-κB and NFAT. Notably, ROS production was enhanced in SHIP-1-deficient GM-BM treated with heat-killed C. albicans, live C. albicans, or the specific Dectin-1 agonists curdlan or whole glucan particles. This increased oxidative burst was dependent on Dectin-1, Syk, PI3K, phosphoinositide-dependent protein kinase 1, and NADPH oxidase. GM-BM from CD11c∆SHIP-1 mice also showed increased killing activity against live C. albicans that was dependent on Dectin-1, Syk, and NADPH oxidase. These results illustrate the complexity of myeloid C-type lectin receptor signaling, and how an activating hemITAM can also couple to intracellular inositol phosphatases to modulate selected functional responses and tightly regulate processes such as ROS production that could be deleterious to the host.
publishDate 2015
dc.date.none.fl_str_mv 2015
2015-11-01
2015
2015-11-01
2020
2020-04-23
dc.type.none.fl_str_mv journal article
http://purl.org/coar/resource_type/c_6501
AM
http://purl.org/coar/version/c_ab4af688f83e57aa
dc.type.openaire.fl_str_mv info:eu-repo/semantics/article
format article
dc.identifier.none.fl_str_mv http://hdl.handle.net/20.500.12105/9706
url http://hdl.handle.net/20.500.12105/9706
dc.language.none.fl_str_mv Inglés
eng
language_invalid_str_mv Inglés
language eng
dc.rights.none.fl_str_mv open access
http://purl.org/coar/access_right/c_abf2
Attribution-NonCommercial-NoDerivatives 4.0 Internacional
http://creativecommons.org/licenses/by-nc-nd/4.0/
dc.rights.openaire.fl_str_mv info:eu-repo/semantics/openAccess
rights_invalid_str_mv open access
http://purl.org/coar/access_right/c_abf2
Attribution-NonCommercial-NoDerivatives 4.0 Internacional
http://creativecommons.org/licenses/by-nc-nd/4.0/
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
dc.publisher.none.fl_str_mv American Association of Immunologists (AAI)
publisher.none.fl_str_mv American Association of Immunologists (AAI)
dc.source.none.fl_str_mv reponame:Repisalud
instname:Instituto de Salud Carlos III (ISCIII)
instname_str Instituto de Salud Carlos III (ISCIII)
reponame_str Repisalud
collection Repisalud
repository.name.fl_str_mv
repository.mail.fl_str_mv
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