Activation of CK1ɛ by PP2A/PR61ɛ is required for the initiation of Wnt signaling

Canonical Wnt signaling induces the stabilization of β-catenin, its translocation to the nucleus and the activation of target promoters. This pathway is initiated by the binding of Wnt ligands to the Frizzled receptor, the association of the LRP5/6 co-receptor and the formation of a complex comprisi...

Full description

Bibliographic Details
Authors: Vinyoles, Meritxell, Valle Pérez, Beatriz del, Curto, Josué, Padilla, Mary, Villarroel, Aida, Yang, J.Y., García de Herreros, Antonio, Duñach, Mireia
Format: article
Status:Versión aceptada para publicación
Publication Date:2017
Country:España
Institution:Universitat Pompeu Fabra
Repository:Repositorio Digital de la UPF
OAI Identifier:oai:repositori.upf.edu:10230/35913
Online Access:http://hdl.handle.net/10230/35913
http://dx.doi.org/10.1038/onc.2016.209
Access Level:Open access
Keyword:Fosfoproteïnes fosfatases
Interacció cel·lular
Description
Summary:Canonical Wnt signaling induces the stabilization of β-catenin, its translocation to the nucleus and the activation of target promoters. This pathway is initiated by the binding of Wnt ligands to the Frizzled receptor, the association of the LRP5/6 co-receptor and the formation of a complex comprising Dvl-2, Axin and protein kinases CK1α, ɛ, γ and GSK3. Among these, activation of CK1ɛ, constitutively bound to LRP5/6 through p120-catenin, is required for the association of the rest of the components. We describe here that CK1ɛ is activated by the PP2A/PR61ɛ phosphatase. Binding of Wnt ligands promotes the interaction of LRP5/6-associated CK1ɛ with Frizzled-bound PR61ɛ regulatory subunit, facilitating the access of PP2A catalytic subunit to CK1ɛ and its activation, what enables the recruitment of Dvl-2 to the receptor complex and the initiation of the Wnt pathway. Our results uncover the mechanism of activation of the canonical Wnt pathway by its ligands.