A gene regulatory network critical for axillary bud dormancy directly controlled by Arabidopsis BRANCHED1

The Arabidopsis thaliana transcription factor BRANCHED1 (BRC1) plays a pivotal role in the control of shoot branching as it integrates environmental and endogenous signals that influence axillary bud growth. Despite its remarkable activity as a growth inhibitor, the mechanisms by which BRC1 promotes...

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Detalles Bibliográficos
Autores: van Es, Sam W, Muñoz-Gasca, Aitor, Romero-Campero, Francisco J, González-Grandío, Eduardo, Reyes, Pedro de los, Tarancón, Carlos, van Dijk, Aalt D J, van Esse, Wilma, Pascual-García, Alberto, Angenent, Gerco C, Immink, Richard G H, Cubas, Pilar
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2024
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/354846
Acceso en línea:http://hdl.handle.net/10261/354846
https://api.elsevier.com/content/abstract/scopus_id/85177892527
Access Level:acceso abierto
Palabra clave:BRANCHED1
Arabidopsis
TCP
Axillary bud dormancy
Gene regulatory network
Shoot branching
Systems biology
Transcription factors
Descripción
Sumario:The Arabidopsis thaliana transcription factor BRANCHED1 (BRC1) plays a pivotal role in the control of shoot branching as it integrates environmental and endogenous signals that influence axillary bud growth. Despite its remarkable activity as a growth inhibitor, the mechanisms by which BRC1 promotes bud dormancy are largely unknown. We determined the genome-wide BRC1 binding sites in vivo and combined these with transcriptomic data and gene co-expression analyses to identify bona fide BRC1 direct targets. Next, we integrated multi-omics data to infer the BRC1 gene regulatory network (GRN) and used graph theory techniques to find network motifs that control the GRN dynamics. We generated an open online tool to interrogate this network. A group of BRC1 target genes encoding transcription factors (BTFs) orchestrate this intricate transcriptional network enriched in abscisic acid-related components. Promoter::β-GLUCURONIDASE transgenic lines confirmed that BTFs are expressed in axillary buds. Transient co-expression assays and studies in planta using mutant lines validated the role of BTFs in modulating the GRN and promoting bud dormancy. This knowledge provides access to the developmental mechanisms that regulate shoot branching and helps identify candidate genes to use as tools to adapt plant architecture and crop production to ever-changing environmental conditions.