Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers

[Significance] Theory and experiments suggest that rising soil nitrogen (N) could delay the cessation of the growing season based on leaf phenology data, but no intercontinental analyses on xylem phenology have been carried out. Using data on xylem phenology from conifers across the Northern Hemisph...

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Autores: Zhang, Yaling, Huang, Jian-Guo, Wang, Minhuang, Wang, Wenjin, Yang, Feiyu, Deslauriers, Annie, Fonti, Patrick, Liang, Eryuan, Mäkinen, Harri, Oberhuber, Walter, Rathgeber, Cyrille, Tognetti, Roberto, Treml, Václav, Yang, Bao, Zhai, Lihong, Antonucci, Serena, Buttò, Valentina, Camarero, Jesús Julio, Campelo, Filipe, Čufar, Katarina, Luis, Martin De, Fajstavr, Marek, Giovannelli, Alessio, Gričar, Jožica, Gruber, Andreas, Gryc, Vladimír, Güney, Aylin, Jyske, Tuula, Kašpar, Jakub, King, Gregory, Krause, Cornelia, Lemay, Audrey, Lombardi, Fabio, Martínez del Castillo, Edurne, Morin, Hubert, Nabais, Cristina, Nöjd, Pekka, Peters, Richard L., Prislan, Peter, Saracino, Antonio, Shishov, Vladimir V., Vavrčík, Hanuš, Vieira, Joana, Zeng, Qiao, Rossi, Sergio
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2025
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/398705
Acceso en línea:http://hdl.handle.net/10261/398705
https://api.elsevier.com/content/abstract/scopus_id/105012093481
Access Level:acceso abierto
Palabra clave:Autumn phenology
Soil moisture
Stem growth
Wood formation
Xylogenesis
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dc.title.none.fl_str_mv Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
title Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
spellingShingle Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
Zhang, Yaling
Autumn phenology
Soil moisture
Stem growth
Wood formation
Xylogenesis
title_short Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
title_full Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
title_fullStr Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
title_full_unstemmed Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
title_sort Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifers
dc.creator.none.fl_str_mv Zhang, Yaling
Huang, Jian-Guo
Wang, Minhuang
Wang, Wenjin
Yang, Feiyu
Deslauriers, Annie
Fonti, Patrick
Liang, Eryuan
Mäkinen, Harri
Oberhuber, Walter
Rathgeber, Cyrille
Tognetti, Roberto
Treml, Václav
Yang, Bao
Zhai, Lihong
Antonucci, Serena
Buttò, Valentina
Camarero, Jesús Julio
Campelo, Filipe
Čufar, Katarina
Luis, Martin De
Fajstavr, Marek
Giovannelli, Alessio
Gričar, Jožica
Gruber, Andreas
Gryc, Vladimír
Güney, Aylin
Jyske, Tuula
Kašpar, Jakub
King, Gregory
Krause, Cornelia
Lemay, Audrey
Lombardi, Fabio
Martínez del Castillo, Edurne
Morin, Hubert
Nabais, Cristina
Nöjd, Pekka
Peters, Richard L.
Prislan, Peter
Saracino, Antonio
Shishov, Vladimir V.
Vavrčík, Hanuš
Vieira, Joana
Zeng, Qiao
Rossi, Sergio
author Zhang, Yaling
author_facet Zhang, Yaling
Huang, Jian-Guo
Wang, Minhuang
Wang, Wenjin
Yang, Feiyu
Deslauriers, Annie
Fonti, Patrick
Liang, Eryuan
Mäkinen, Harri
Oberhuber, Walter
Rathgeber, Cyrille
Tognetti, Roberto
Treml, Václav
Yang, Bao
Zhai, Lihong
Antonucci, Serena
Buttò, Valentina
Camarero, Jesús Julio
Campelo, Filipe
Čufar, Katarina
Luis, Martin De
Fajstavr, Marek
Giovannelli, Alessio
Gričar, Jožica
Gruber, Andreas
Gryc, Vladimír
Güney, Aylin
Jyske, Tuula
Kašpar, Jakub
King, Gregory
Krause, Cornelia
Lemay, Audrey
Lombardi, Fabio
Martínez del Castillo, Edurne
Morin, Hubert
Nabais, Cristina
Nöjd, Pekka
Peters, Richard L.
Prislan, Peter
Saracino, Antonio
Shishov, Vladimir V.
Vavrčík, Hanuš
Vieira, Joana
Zeng, Qiao
Rossi, Sergio
author_role author
author2 Huang, Jian-Guo
Wang, Minhuang
Wang, Wenjin
Yang, Feiyu
Deslauriers, Annie
Fonti, Patrick
Liang, Eryuan
Mäkinen, Harri
Oberhuber, Walter
Rathgeber, Cyrille
Tognetti, Roberto
Treml, Václav
Yang, Bao
Zhai, Lihong
Antonucci, Serena
Buttò, Valentina
Camarero, Jesús Julio
Campelo, Filipe
Čufar, Katarina
Luis, Martin De
Fajstavr, Marek
Giovannelli, Alessio
Gričar, Jožica
Gruber, Andreas
Gryc, Vladimír
Güney, Aylin
Jyske, Tuula
Kašpar, Jakub
King, Gregory
Krause, Cornelia
Lemay, Audrey
Lombardi, Fabio
Martínez del Castillo, Edurne
Morin, Hubert
Nabais, Cristina
Nöjd, Pekka
Peters, Richard L.
Prislan, Peter
Saracino, Antonio
Shishov, Vladimir V.
Vavrčík, Hanuš
Vieira, Joana
Zeng, Qiao
Rossi, Sergio
author2_role author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
author
dc.contributor.none.fl_str_mv National Natural Science Foundation of China
Zhejiang Province
Xinjiang Uygur Autonomous Region
Zhejiang University
Austrian Science Fund
Consortium de Recherche sur la Forêt Boréale Commerciale (Canada)
Fonds de Recherche Nature et Technologies (Canada)
Forêt d'Enseignement et de Recherche Simoncouche
Observatoire Régional de Recherche sur la Forêt Boréale (Canada)
Ministry of Science and Higher Education of the Russian Federation
Natural Sciences and Engineering Research Council of Canada
Slovenian Research Agency
European Commission
Swiss National Science Foundation
Agence Nationale de la Recherche (France)
Academy of Finland
Technology Agency of the Czech Republic
Provincia Autonoma di Trento
Fondation pour la Recherche sur la Biodiversité
Camarero, Jesús Julio [0000-0003-2436-2922]
Consejo Superior de Investigaciones Científicas [https://ror.org/02gfc7t72]
dc.subject.none.fl_str_mv Autumn phenology
Soil moisture
Stem growth
Wood formation
Xylogenesis
topic Autumn phenology
Soil moisture
Stem growth
Wood formation
Xylogenesis
description [Significance] Theory and experiments suggest that rising soil nitrogen (N) could delay the cessation of the growing season based on leaf phenology data, but no intercontinental analyses on xylem phenology have been carried out. Using data on xylem phenology from conifers across the Northern Hemisphere, we found that rising soil N delays the end of xylem cell wall thickening but advances the cessation of cell enlargement. While xylem cell enlargement is responsible for stem size growth, cell wall thickening accounts for 90% of woody biomass production. The contrasting effects of soil N on these two xylem differentiation processes would affect xylem cell anatomy and consequently influence water transport and wood density in conifers.
publishDate 2025
dc.date.none.fl_str_mv 2025
2025
2025
dc.type.none.fl_str_mv info:eu-repo/semantics/article
http://purl.org/coar/resource_type/c_6501
Publisher's version
info:eu-repo/semantics/publishedVersion
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv http://hdl.handle.net/10261/398705
https://api.elsevier.com/content/abstract/scopus_id/105012093481
url http://hdl.handle.net/10261/398705
https://api.elsevier.com/content/abstract/scopus_id/105012093481
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
dc.relation.none.fl_str_mv #PLACEHOLDER_PARENT_METADATA_VALUE#
info:eu-repo/grantAgreement/EC/H2020/952314
The underlying dataset has been published as supplementary material of the article in the publisher platform at DOI https://doi.org/10.1073/pnas.2421834122
https://doi.org/10.1073/pnas.2421834122

dc.rights.none.fl_str_mv info:eu-repo/semantics/openAccess
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
dc.publisher.none.fl_str_mv National Academy of Sciences (U.S.)
publisher.none.fl_str_mv National Academy of Sciences (U.S.)
dc.source.none.fl_str_mv reponame:DIGITAL.CSIC. Repositorio Institucional del CSIC
instname:Consejo Superior de Investigaciones Científicas (CSIC)
instname_str Consejo Superior de Investigaciones Científicas (CSIC)
reponame_str DIGITAL.CSIC. Repositorio Institucional del CSIC
collection DIGITAL.CSIC. Repositorio Institucional del CSIC
repository.name.fl_str_mv
repository.mail.fl_str_mv
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spelling Soil nitrogen drives inverse acclimation of xylem growth cessation to rising temperature in Northern Hemisphere conifersZhang, YalingHuang, Jian-GuoWang, MinhuangWang, WenjinYang, FeiyuDeslauriers, AnnieFonti, PatrickLiang, EryuanMäkinen, HarriOberhuber, WalterRathgeber, CyrilleTognetti, RobertoTreml, VáclavYang, BaoZhai, LihongAntonucci, SerenaButtò, ValentinaCamarero, Jesús JulioCampelo, FilipeČufar, KatarinaLuis, Martin DeFajstavr, MarekGiovannelli, AlessioGričar, JožicaGruber, AndreasGryc, VladimírGüney, AylinJyske, TuulaKašpar, JakubKing, GregoryKrause, CorneliaLemay, AudreyLombardi, FabioMartínez del Castillo, EdurneMorin, HubertNabais, CristinaNöjd, PekkaPeters, Richard L.Prislan, PeterSaracino, AntonioShishov, Vladimir V.Vavrčík, HanušVieira, JoanaZeng, QiaoRossi, SergioAutumn phenologySoil moistureStem growthWood formationXylogenesis[Significance] Theory and experiments suggest that rising soil nitrogen (N) could delay the cessation of the growing season based on leaf phenology data, but no intercontinental analyses on xylem phenology have been carried out. Using data on xylem phenology from conifers across the Northern Hemisphere, we found that rising soil N delays the end of xylem cell wall thickening but advances the cessation of cell enlargement. While xylem cell enlargement is responsible for stem size growth, cell wall thickening accounts for 90% of woody biomass production. The contrasting effects of soil N on these two xylem differentiation processes would affect xylem cell anatomy and consequently influence water transport and wood density in conifers.[Abstract] Controlled experiments suggest that the seasonal build-up of nitrogen (N) limitation constrains the responses of forest autumn phenology to elevated temperatures. Therefore, rising soil N is expected to increase the delaying effects of elevated temperature on the end of the season, i.e., leaf senescence. However, the interactive effects of temperature, soil N, and aridity on xylem autumn phenology remain unknown. We conducted a wide spatial analysis from 75 conifer sites in the Northern Hemisphere and found that rising soil N increases the delaying effects of elevated temperature on the end of xylem cell wall thickening but reduced the delaying effects on the cessation of cell enlargement, especially in humid regions. The contrasting effects of elevated soil N on cell enlargement versus cell wall thickening could affect xylem cell anatomy, thereby induce changes in wood density, and induce a decoupling of stem size growth from photosynthate production. These analyses extend previous findings on forest autumn phenology by systematically investigating the spatial variation in the interactive effects of temperature and soil N on xylem autumn phenology at the cellular scale.This work was funded by the National Natural Science Foundation of China (Grant Nos. 32271653,42471052, and 32401377), the Key R&D Program of Zhejiang (2024C03243), GDAS’ Project of Science and Technology Development (2024GDASZH-2024010101), the Xinjiang Regional Collaborative Innovation Project (2022E01045), and Zhejiang University (108000*1942222R1). Additional funding came from the Austrian Science Fund (FWF P22280-B16 and P25643-B16), Consortium de Recherche sur la Forêt Boréale Commerciale, Fonds de Recherche sur la Nature et les Technologies du Québec, Forêt d’enseignement et de recherche Simoncouche, Observatoire régional de recherche en forêt boréale, The Ministry of Science and Higher Education of the Russian Federation (project #FSRZ-2023-0007), Natural Sciences and Engineering Research Council of Canada, Slovenian Research and Innovation Agency ARIS (Research Core Funding Nos. P4-0430 and P4-0015, Projects J4-2541, J4-4541, and Z4-7318), and the European Union’s Horizon 2020 research and innovation program under the ASFORCLIC grant agreement (No. 952314). Further support was provided by MIUR-PRIN 2002 (2002075152) and 2005 (2005072877), the Swiss NSF (Projects INTEGRAL-121859 and LOTFOR-150205 and CALEIDOSCOPE-212902), the French National Research Agency (ANR) as part of the “Investissements d’Avenir” program (ANR-11-LABX-0002-01, Lab of Excellence ARBRE), Academy of Finland (Nos. 250299, 257641, and 265504), Grant Agency of Czech Republic (P504/11/P557), and Provincia Autonoma di Trento (Project “SOFIE 2”–3012/2007). CBKR would like to thank the SILVATECH platform (Silvatech, INRAE, 2018. Structural and functional analysis of tree and wood Facility, doi: https://doi.org/10.15454/1.5572400113627854E12) for its contribution to the acquisition of wood formation monitoring data. Cooperation among the authors was supported by the EU COST Action FP1106 STReESS. The views and conclusions expressed in this document are those of the authors and do not necessarily reflect the opinions or policies of the funding agencies or supporting institutions. We would like to acknowledge the FAIRWood project funded by the CESAB of the French Foundation for Research on Biodiversity (FRB), and specially its GitHub repository for providing the R script and some functions that were helpful in processing the data (https://github.com/FAIRWood-datashare/DataProcessingFunctions).Peer reviewedNational Academy of Sciences (U.S.)National Natural Science Foundation of ChinaZhejiang ProvinceXinjiang Uygur Autonomous RegionZhejiang UniversityAustrian Science FundConsortium de Recherche sur la Forêt Boréale Commerciale (Canada)Fonds de Recherche Nature et Technologies (Canada)Forêt d'Enseignement et de Recherche SimoncoucheObservatoire Régional de Recherche sur la Forêt Boréale (Canada)Ministry of Science and Higher Education of the Russian FederationNatural Sciences and Engineering Research Council of CanadaSlovenian Research AgencyEuropean CommissionSwiss National Science FoundationAgence Nationale de la Recherche (France)Academy of FinlandTechnology Agency of the Czech RepublicProvincia Autonoma di TrentoFondation pour la Recherche sur la BiodiversitéCamarero, Jesús Julio [0000-0003-2436-2922]Consejo Superior de Investigaciones Científicas [https://ror.org/02gfc7t72]202520252025info:eu-repo/semantics/articlehttp://purl.org/coar/resource_type/c_6501Publisher's versioninfo:eu-repo/semantics/publishedVersionapplication/pdfhttp://hdl.handle.net/10261/398705https://api.elsevier.com/content/abstract/scopus_id/105012093481reponame:DIGITAL.CSIC. Repositorio Institucional del CSICinstname:Consejo Superior de Investigaciones Científicas (CSIC)Inglés#PLACEHOLDER_PARENT_METADATA_VALUE#info:eu-repo/grantAgreement/EC/H2020/952314The underlying dataset has been published as supplementary material of the article in the publisher platform at DOI https://doi.org/10.1073/pnas.2421834122https://doi.org/10.1073/pnas.2421834122Síinfo:eu-repo/semantics/openAccessoai:digital.csic.es:10261/3987052026-05-22T06:33:51Z
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