Interaction Dynamics Determine Signaling and Output Pathway Responses

The understanding of interaction dynamics in signaling pathways can shed light on pathway architecture and provide insights into targets for intervention. Here, we explored the relevance of kinetic rate constants of a key upstream osmosensor in the yeast high-osmolarity glycerol-mitogen-activated pr...

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Detalles Bibliográficos
Autores: Stojanovski, Klement, Ferrar, Tony, Benisty, Hannah, 1986-, Uschner, Friedemann, Delgado Blanco, Javier, Jimenez, Javier, Solé, Carme, Nadal Clanchet, Eulàlia de, Klipp, E., 1965-, Posas Garriga, Francesc, Serrano Pubull, Luis, 1982-, Kiel, Christina
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2017
País:España
Institución:Varias* (Consorci de Biblioteques Universitáries de Catalunya, Centre de Serveis Científics i Acadèmics de Catalunya)
Repositorio:Recercat. Dipósit de la Recerca de Catalunya
OAI Identifier:oai:recercat.cat:10230/32109
Acceso en línea:http://hdl.handle.net/10230/32109
http://dx.doi.org/10.1016/j.celrep.2017.03.029
Access Level:acceso abierto
Palabra clave:Kinetic perturbations
Phosphorelay
HOG-MAPK pathway
Osmostress response
Descripción
Sumario:The understanding of interaction dynamics in signaling pathways can shed light on pathway architecture and provide insights into targets for intervention. Here, we explored the relevance of kinetic rate constants of a key upstream osmosensor in the yeast high-osmolarity glycerol-mitogen-activated protein kinase (HOG-MAPK) pathway to signaling output responses. We created mutant pairs of the Sln1-Ypd1 complex interface that caused major compensating changes in the association (kon) and dissociation (koff) rate constants (kinetic perturbations) but only moderate changes in the overall complex affinity (Kd). Yeast cells carrying a Sln1-Ypd1 mutant pair with moderate increases in kon and koff displayed a lower threshold of HOG pathway activation than wild-type cells. Mutants with higher kon and koff rates gave rise to higher basal signaling and gene expression but impaired osmoadaptation. Thus, the kon and koff rates of the components in the Sln1 osmosensor determine proper signaling dynamics and osmoadaptation.