High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary

Subterranean estuaries are biogeochemically active coastal sites resulting from the underground mixing of fresh aquifer groundwater and seawater. In these systems, microbial activity can largely transform the chemical elements that may reach the sea through submarine groundwater discharge (SGD), but...

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Autores: Ruiz González, Clara, Rodríguez Pie, Lara, Maister, Olena, Rodellas, Valentí, Alorda Keinglass, Aaron, Diego Feliu, Marc|||0000-0002-4782-3210, Folch Sancho, Albert|||0000-0002-8490-1038, Garcia Orellana, Jordi, Gasol, Josep Maria
Tipo de recurso: artículo
Fecha de publicación:2022
País:España
Institución:Universitat Politècnica de Catalunya (UPC)
Repositorio:UPCommons. Portal del coneixement obert de la UPC
Idioma:inglés
OAI Identifier:oai:upcommons.upc.edu:2117/377668
Acceso en línea:https://hdl.handle.net/2117/377668
https://dx.doi.org/10.1111/mec.16695
Access Level:acceso abierto
Palabra clave:Groundwater
Aquatic prokaryotic communities
Coastal aquifer
Microbial activity
Mixing zone
Patescibacteria
Seawater intrusion
Submarine groundwater discharge
Subterranean estuaries
Terrestrial–marine interface
Aigües subterrànies
Àrees temàtiques de la UPC::Enginyeria civil::Geologia::Hidrologia subterrània
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network_acronym_str ES
network_name_str España
repository_id_str
dc.title.none.fl_str_mv High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
title High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
spellingShingle High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
Ruiz González, Clara
Groundwater
Aquatic prokaryotic communities
Coastal aquifer
Microbial activity
Mixing zone
Patescibacteria
Seawater intrusion
Submarine groundwater discharge
Subterranean estuaries
Terrestrial–marine interface
Aigües subterrànies
Àrees temàtiques de la UPC::Enginyeria civil::Geologia::Hidrologia subterrània
title_short High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
title_full High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
title_fullStr High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
title_full_unstemmed High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
title_sort High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuary
dc.creator.none.fl_str_mv Ruiz González, Clara
Rodríguez Pie, Lara
Maister, Olena
Rodellas, Valentí
Alorda Keinglass, Aaron
Diego Feliu, Marc|||0000-0002-4782-3210
Folch Sancho, Albert|||0000-0002-8490-1038
Garcia Orellana, Jordi
Gasol, Josep Maria
author Ruiz González, Clara
author_facet Ruiz González, Clara
Rodríguez Pie, Lara
Maister, Olena
Rodellas, Valentí
Alorda Keinglass, Aaron
Diego Feliu, Marc|||0000-0002-4782-3210
Folch Sancho, Albert|||0000-0002-8490-1038
Garcia Orellana, Jordi
Gasol, Josep Maria
author_role author
author2 Rodríguez Pie, Lara
Maister, Olena
Rodellas, Valentí
Alorda Keinglass, Aaron
Diego Feliu, Marc|||0000-0002-4782-3210
Folch Sancho, Albert|||0000-0002-8490-1038
Garcia Orellana, Jordi
Gasol, Josep Maria
author2_role author
author
author
author
author
author
author
author
dc.subject.none.fl_str_mv Groundwater
Aquatic prokaryotic communities
Coastal aquifer
Microbial activity
Mixing zone
Patescibacteria
Seawater intrusion
Submarine groundwater discharge
Subterranean estuaries
Terrestrial–marine interface
Aigües subterrànies
Àrees temàtiques de la UPC::Enginyeria civil::Geologia::Hidrologia subterrània
topic Groundwater
Aquatic prokaryotic communities
Coastal aquifer
Microbial activity
Mixing zone
Patescibacteria
Seawater intrusion
Submarine groundwater discharge
Subterranean estuaries
Terrestrial–marine interface
Aigües subterrànies
Àrees temàtiques de la UPC::Enginyeria civil::Geologia::Hidrologia subterrània
description Subterranean estuaries are biogeochemically active coastal sites resulting from the underground mixing of fresh aquifer groundwater and seawater. In these systems, microbial activity can largely transform the chemical elements that may reach the sea through submarine groundwater discharge (SGD), but little is known about the microorganisms thriving in these land-sea transition zones. We present the first spatially-resolved characterization of the bacterial assemblages along a coastal aquifer in the NW Mediterranean, considering the entire subsurface salinity gradient. Combining bulk heterotrophic activity measurements, flow cytometry, microscopy and 16S rRNA gene sequencing we find large variations in prokaryotic abundances, cell size, activity and diversity at both the horizontal and vertical scales that reflect the pronounced physicochemical gradients. The parts of the transect most influenced by freshwater were characterized by smaller cells and lower prokaryotic abundances and heterotrophic production, but some activity hotspots were found at deep low-oxygen saline groundwater sites enriched in nitrite and ammonium. Diverse, heterogeneous and highly endemic communities dominated by Proteobacteria, Patescibacteria, Desulfobacterota and Bacteroidota were observed throughout the aquifer, pointing to clearly differentiated prokaryotic niches across these transition zones and little microbial connectivity between groundwater and Mediterranean seawater habitats. Finally, experimental manipulations unveiled large increases in community heterotrophic activity driven by fast growth of some rare and site-specific groundwater Proteobacteria. Our results indicate that prokaryotic communities within subterranean estuaries are highly heterogeneous in terms of biomass, activity and diversity, suggesting that their role in transforming nutrients will also vary spatially within these terrestrial–marine transition zones.
publishDate 2022
dc.date.none.fl_str_mv 2022
2022-11-01
2022
2022-12-05
dc.type.none.fl_str_mv journal article
http://purl.org/coar/resource_type/c_6501
VoR
http://purl.org/coar/version/c_970fb48d4fbd8a85
dc.type.openaire.fl_str_mv info:eu-repo/semantics/article
format article
dc.identifier.none.fl_str_mv https://hdl.handle.net/2117/377668
https://dx.doi.org/10.1111/mec.16695
url https://hdl.handle.net/2117/377668
https://dx.doi.org/10.1111/mec.16695
dc.language.none.fl_str_mv Inglés
eng
language_invalid_str_mv Inglés
language eng
dc.rights.none.fl_str_mv open access
http://purl.org/coar/access_right/c_abf2
Attribution-NonCommercial-NoDerivatives 4.0 International
http://creativecommons.org/licenses/by-nc-nd/4.0/
dc.rights.openaire.fl_str_mv info:eu-repo/semantics/openAccess
rights_invalid_str_mv open access
http://purl.org/coar/access_right/c_abf2
Attribution-NonCommercial-NoDerivatives 4.0 International
http://creativecommons.org/licenses/by-nc-nd/4.0/
eu_rights_str_mv openAccess
dc.format.none.fl_str_mv application/pdf
dc.source.none.fl_str_mv reponame:UPCommons. Portal del coneixement obert de la UPC
instname:Universitat Politècnica de Catalunya (UPC)
instname_str Universitat Politècnica de Catalunya (UPC)
reponame_str UPCommons. Portal del coneixement obert de la UPC
collection UPCommons. Portal del coneixement obert de la UPC
repository.name.fl_str_mv
repository.mail.fl_str_mv
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spelling High spatial heterogeneity and low connectivity of bacterial communities along a Mediterranean subterranean estuaryRuiz González, ClaraRodríguez Pie, LaraMaister, OlenaRodellas, ValentíAlorda Keinglass, AaronDiego Feliu, Marc|||0000-0002-4782-3210Folch Sancho, Albert|||0000-0002-8490-1038Garcia Orellana, JordiGasol, Josep MariaGroundwaterAquatic prokaryotic communitiesCoastal aquiferMicrobial activityMixing zonePatescibacteriaSeawater intrusionSubmarine groundwater dischargeSubterranean estuariesTerrestrial–marine interfaceAigües subterràniesÀrees temàtiques de la UPC::Enginyeria civil::Geologia::Hidrologia subterràniaSubterranean estuaries are biogeochemically active coastal sites resulting from the underground mixing of fresh aquifer groundwater and seawater. In these systems, microbial activity can largely transform the chemical elements that may reach the sea through submarine groundwater discharge (SGD), but little is known about the microorganisms thriving in these land-sea transition zones. We present the first spatially-resolved characterization of the bacterial assemblages along a coastal aquifer in the NW Mediterranean, considering the entire subsurface salinity gradient. Combining bulk heterotrophic activity measurements, flow cytometry, microscopy and 16S rRNA gene sequencing we find large variations in prokaryotic abundances, cell size, activity and diversity at both the horizontal and vertical scales that reflect the pronounced physicochemical gradients. The parts of the transect most influenced by freshwater were characterized by smaller cells and lower prokaryotic abundances and heterotrophic production, but some activity hotspots were found at deep low-oxygen saline groundwater sites enriched in nitrite and ammonium. Diverse, heterogeneous and highly endemic communities dominated by Proteobacteria, Patescibacteria, Desulfobacterota and Bacteroidota were observed throughout the aquifer, pointing to clearly differentiated prokaryotic niches across these transition zones and little microbial connectivity between groundwater and Mediterranean seawater habitats. Finally, experimental manipulations unveiled large increases in community heterotrophic activity driven by fast growth of some rare and site-specific groundwater Proteobacteria. Our results indicate that prokaryotic communities within subterranean estuaries are highly heterogeneous in terms of biomass, activity and diversity, suggesting that their role in transforming nutrients will also vary spatially within these terrestrial–marine transition zones.This work was funded by the Spanish Ministry of Science, Innovation and Universities (MICINN) through the GRAMMI project (RTI2018-099740-J-I00), the Ramon y Cajal contract (RYC2019-026758-I) and partly by the project MEDISTRAES III (PID2019-110311RB-C22 and PID2019-110212RB-C21). Additional financial support was provided by grants SPIP2020-02595 (Ministry for the Ecological Transition and the Demographic Challenge [MITECO]) with funding from the Spanish government through the ‘Severo Ochoa Centre of Excellence’ accreditation (CEX2019-000928-S). JGO acknowledges the financial support of the Spanish Ministry of Science, Innovation and Universities, through the “Maria de Maeztu” programme for Units of Excellence (CEX2019-000940-M). VR acknowledges financial support from the Beatriu de Pinós postdoctoral programme of the Catalan Government (2019-BP-00241). AAK acknowledges financial support from ICTA “Unit of Excellence” (MinECo, MDM2015-0552-17-1) and PhD fellowship, BES-2017-080740. The coauthor AF is a Serra Húnter Fellow. MDF acknowledges the economic support from the FI-2017 fellowships of the Generalitat de Catalunya autonomous government (2017FI_B_00365). The authors would like to thank the support of the Generalitat de Catalunya to MERS (2017 SGR-1588), GHS (2017 SGR 1485) and 2017SGR/156, for additional funding. We would like to thank SIMMAR (Serveis Integrals de Manteniment del Maresme) and the Consell Comarcal del Maresme in the construction of the research site. A special word for Jordi Garcia-Orellana, great mentor and friend, who left us recently and unexpectedly, and without whom this and much of the ongoing work would not exist. May he rest in peace.Peer Reviewed20222022-11-0120222022-12-05journal articlehttp://purl.org/coar/resource_type/c_6501VoRhttp://purl.org/coar/version/c_970fb48d4fbd8a85info:eu-repo/semantics/articleapplication/pdfhttps://hdl.handle.net/2117/377668https://dx.doi.org/10.1111/mec.16695reponame:UPCommons. Portal del coneixement obert de la UPCinstname:Universitat Politècnica de Catalunya (UPC)Inglésengopen accesshttp://purl.org/coar/access_right/c_abf2Attribution-NonCommercial-NoDerivatives 4.0 Internationalhttp://creativecommons.org/licenses/by-nc-nd/4.0/info:eu-repo/semantics/openAccessoai:upcommons.upc.edu:2117/3776682026-05-27T15:37:01Z
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