Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species

An improved definition of tree vigor is required to characterize forest resilience in response to drought. We developed a vigor index based on annual crown defoliation and radial-growth data. The index was applied to stone pine (Pinus pinea) and cork oak (Quercus suber) trees affected by die-off in...

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Autores: Camarero, Jesús Julio, Gazol Burgos, Antonio, Díaz-Delgado, Ricardo, Aragonés, David, Valeriano, Cristina, Fernández Cortés, Ángel
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2025
País:España
Institución:Consejo Superior de Investigaciones Científicas (CSIC)
Repositorio:DIGITAL.CSIC. Repositorio Institucional del CSIC
OAI Identifier:oai:digital.csic.es:10261/403145
Acceso en línea:http://hdl.handle.net/10261/403145
https://api.elsevier.com/content/abstract/scopus_id/105012245653
Access Level:acceso abierto
Palabra clave:Crown defoliation
Dendrometer
Drought stress
Forest die-off
Growth decline
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dc.title.none.fl_str_mv Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
title Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
spellingShingle Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
Camarero, Jesús Julio
Crown defoliation
Dendrometer
Drought stress
Forest die-off
Growth decline
title_short Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
title_full Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
title_fullStr Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
title_full_unstemmed Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
title_sort Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak species
dc.creator.none.fl_str_mv Camarero, Jesús Julio
Gazol Burgos, Antonio
Díaz-Delgado, Ricardo
Aragonés, David
Valeriano, Cristina
Fernández Cortés, Ángel
author Camarero, Jesús Julio
author_facet Camarero, Jesús Julio
Gazol Burgos, Antonio
Díaz-Delgado, Ricardo
Aragonés, David
Valeriano, Cristina
Fernández Cortés, Ángel
author_role author
author2 Gazol Burgos, Antonio
Díaz-Delgado, Ricardo
Aragonés, David
Valeriano, Cristina
Fernández Cortés, Ángel
author2_role author
author
author
author
author
dc.contributor.none.fl_str_mv National Science Foundation (US)
Department of Agriculture (US)
Department of Energy (US)
U.S. Geological Survey
Ministerio de Ciencia, Innovación y Universidades (España)
European Commission
Consejo Superior de Investigaciones Científicas (España)
Agencia Estatal de Investigación (España)
Camarero, Jesús Julio [0000-0003-2436-2922]
Gazol Burgos, Antonio [0000-0001-5902-9543]
Díaz-Delgado, Ricardo [0000-0002-0460-4616]
Aragonés, David [0000-0003-4989-9005]
Valeriano, Cristina [0000-0001-7687-1417]
Consejo Superior de Investigaciones Científicas [https://ror.org/02gfc7t72]
dc.subject.none.fl_str_mv Crown defoliation
Dendrometer
Drought stress
Forest die-off
Growth decline
topic Crown defoliation
Dendrometer
Drought stress
Forest die-off
Growth decline
description An improved definition of tree vigor is required to characterize forest resilience in response to drought. We developed a vigor index based on annual crown defoliation and radial-growth data. The index was applied to stone pine (Pinus pinea) and cork oak (Quercus suber) trees affected by die-off in the Doñana National Park (SW Spain), comparing a dry and a wet site. Climate data were correlated with defoliation, growth and the vigor index. Daily measures of cover greenness (gcc, green chromatic coordinate), obtained from phenocams (period 2018 −2024), and changes in stem radial growth and tree water deficit, obtained from dendrometers (2024 growing season), were also investigated. Finally, we used δ13C wood data to retrospectively assess intrinsic water-use efficiency (iWUE). During 2024, pines grew more than oaks on a yearly basis according to dendrometer data (2.0 vs. 0.5 mm), but annual tree rings were wider in oaks (4.34 mm) than in pines (2.50 mm). Tree water deficit was higher in oaks. In the dry site, gcc and stone pine water deficit were negatively associated, whereas in the wet site gcc and the cork oak growth rate were positively associated. Cork oak showed higher annual defoliation (26 %) than the pine (22 %). Defoliation increased in both species, whilst basal area increment decreased after the 2005 and 2012 droughts in stone pine and cork oak, respectively. The vigor index declined in the stone pine. Dry and warm winter-to-spring conditions and low soil moisture levels increased defoliation and reduced growth. Cork oak and stone pine responded more to drought in terms of defoliation and growth, respectively. The stone pine showed a sharp increase of iWUE after 2016, but the oak presented a more recent iWUE enhancement. Future research can refine the vigor index using longer time series of defoliation and growth data from multiple tree species.
publishDate 2025
dc.date.none.fl_str_mv 2025
2025
2025
dc.type.none.fl_str_mv info:eu-repo/semantics/article
http://purl.org/coar/resource_type/c_6501
Publisher's version
info:eu-repo/semantics/publishedVersion
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv http://hdl.handle.net/10261/403145
https://api.elsevier.com/content/abstract/scopus_id/105012245653
url http://hdl.handle.net/10261/403145
https://api.elsevier.com/content/abstract/scopus_id/105012245653
dc.language.none.fl_str_mv Inglés
language_invalid_str_mv Inglés
dc.relation.none.fl_str_mv #PLACEHOLDER_PARENT_METADATA_VALUE#
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info:eu-repo/grantAgreement/AEI/Plan Estatal de Investigación Científica y Técnica y de Innovación 2021-2023/PID2021-123675OB-C43
info:eu-repo/grantAgreement/AEI//TED2021–129770B-C21
The underlying dataset has been published as supplementary material of the article in the publisher platform at DOI https://doi.org/10.1016/j.foreco.2025.123050
https://doi.org/10.1016/j.foreco.2025.123050

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dc.publisher.none.fl_str_mv Elsevier BV
publisher.none.fl_str_mv Elsevier BV
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spelling Towards a tree vigor definition: Different responses of defoliation, growth and water-use efficiency to drought in pine and oak speciesCamarero, Jesús JulioGazol Burgos, AntonioDíaz-Delgado, RicardoAragonés, DavidValeriano, CristinaFernández Cortés, ÁngelCrown defoliationDendrometerDrought stressForest die-offGrowth declineAn improved definition of tree vigor is required to characterize forest resilience in response to drought. We developed a vigor index based on annual crown defoliation and radial-growth data. The index was applied to stone pine (Pinus pinea) and cork oak (Quercus suber) trees affected by die-off in the Doñana National Park (SW Spain), comparing a dry and a wet site. Climate data were correlated with defoliation, growth and the vigor index. Daily measures of cover greenness (gcc, green chromatic coordinate), obtained from phenocams (period 2018 −2024), and changes in stem radial growth and tree water deficit, obtained from dendrometers (2024 growing season), were also investigated. Finally, we used δ13C wood data to retrospectively assess intrinsic water-use efficiency (iWUE). During 2024, pines grew more than oaks on a yearly basis according to dendrometer data (2.0 vs. 0.5 mm), but annual tree rings were wider in oaks (4.34 mm) than in pines (2.50 mm). Tree water deficit was higher in oaks. In the dry site, gcc and stone pine water deficit were negatively associated, whereas in the wet site gcc and the cork oak growth rate were positively associated. Cork oak showed higher annual defoliation (26 %) than the pine (22 %). Defoliation increased in both species, whilst basal area increment decreased after the 2005 and 2012 droughts in stone pine and cork oak, respectively. The vigor index declined in the stone pine. Dry and warm winter-to-spring conditions and low soil moisture levels increased defoliation and reduced growth. Cork oak and stone pine responded more to drought in terms of defoliation and growth, respectively. The stone pine showed a sharp increase of iWUE after 2016, but the oak presented a more recent iWUE enhancement. Future research can refine the vigor index using longer time series of defoliation and growth data from multiple tree species.Data used in this research were provided by the PhenoCam Network, which has been supported by the National Science Foundation, the Long-Term Agroecosystem Research (LTAR) network which is supported by the United States Department of Agriculture (USDA), the U.S. Department of Energy, the U.S. Geological Survey, the Northeastern States Research Cooperative, and the USA National Phenology Network. We thank the PhenoCam Network collaborators, including site PIs and technicians, for publicly sharing the data that were used in this paper. Phenocam sites are maintained by the ICTS Doñana Scientific Reserve, funded by the Spanish Ministry of Science. The authors thank ICTS-RBD (Infraestructura Científica y Técnica Singular- Reserva Biológica de Doñana) funded by ICTS-MICIN program, and the managers of DBR for their help in accessing sampled sites. Phenocams were acquired through the project “Adaptation and improvement of the ICTS e-Infrastructure ICTS-RBD for LifeWatch (FEDER FICTS-2014/01/AIC-A-2011–0706)” through the European Regional Development Fund [SUMHAL, LIFEWATCH-2019–09-CSIC-4, POPE 2014–2020] and eLTER Plus project (INFRAIA, Horizon 2020, Agreement no. 871128). This study was funded by projects PID2021–123675OB-C43 and TED2021–129770B-C21 (Spanish Ministry of Science and Innovation). AG was supported by by CSIC under grant PIE-20223AT003.Peer reviewedElsevier BVNational Science Foundation (US)Department of Agriculture (US)Department of Energy (US)U.S. Geological SurveyMinisterio de Ciencia, Innovación y Universidades (España)European CommissionConsejo Superior de Investigaciones Científicas (España)Agencia Estatal de Investigación (España)Camarero, Jesús Julio [0000-0003-2436-2922]Gazol Burgos, Antonio [0000-0001-5902-9543]Díaz-Delgado, Ricardo [0000-0002-0460-4616]Aragonés, David [0000-0003-4989-9005]Valeriano, Cristina [0000-0001-7687-1417]Consejo Superior de Investigaciones Científicas [https://ror.org/02gfc7t72]202520252025info:eu-repo/semantics/articlehttp://purl.org/coar/resource_type/c_6501Publisher's versioninfo:eu-repo/semantics/publishedVersionapplication/pdfhttp://hdl.handle.net/10261/403145https://api.elsevier.com/content/abstract/scopus_id/105012245653reponame:DIGITAL.CSIC. Repositorio Institucional del CSICinstname:Consejo Superior de Investigaciones Científicas (CSIC)Inglés#PLACEHOLDER_PARENT_METADATA_VALUE##PLACEHOLDER_PARENT_METADATA_VALUE#info:eu-repo/grantAgreement/AEI/Plan Estatal de Investigación Científica y Técnica y de Innovación 2021-2023/PID2021-123675OB-C43info:eu-repo/grantAgreement/AEI//TED2021–129770B-C21The underlying dataset has been published as supplementary material of the article in the publisher platform at DOI https://doi.org/10.1016/j.foreco.2025.123050https://doi.org/10.1016/j.foreco.2025.123050Síinfo:eu-repo/semantics/openAccessoai:digital.csic.es:10261/4031452026-05-22T06:33:51Z
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