An evolutionary analysis of the reaction mechanisms of photosystem I reduction by cytochrome c6 and plastocyanin

Photosystem I reduction by the soluble metalloproteins cytochrome c6 and plastocyanin, which are alternatively synthesized by some photosynthetic organisms depending on the relative availability of copper and iron, has been investigated in cyanobacteria, green algae and plants. The reaction mechanis...

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Detalles Bibliográficos
Autores: Rosa Acosta, Miguel Ángel de la, Navarro Carruesco, José Antonio, Díaz Quintana, Antonio Jesús, Cerda Haynes, Berta de la, Molina Heredia, Fernando Publio, Balme, Alexis, Murdoch, Piedad del Socorro, Díaz Moreno, Irene, Durán Díaz, Raúl V., Hervás Morón, Manuel
Tipo de recurso: artículo
Estado:Versión aceptada para publicación
Fecha de publicación:2002
País:España
Institución:Universidad de Sevilla (US)
Repositorio:idUS. Depósito de Investigación de la Universidad de Sevilla
OAI Identifier:oai:idus.us.es:11441/84934
Acceso en línea:https://hdl.handle.net/11441/84934
https://doi.org/10.1016/S1567-5394(01)00136-0
Access Level:acceso abierto
Palabra clave:Cytochrome c6
Plastocyanin
Photosystem
Electron transfer
Descripción
Sumario:Photosystem I reduction by the soluble metalloproteins cytochrome c6 and plastocyanin, which are alternatively synthesized by some photosynthetic organisms depending on the relative availability of copper and iron, has been investigated in cyanobacteria, green algae and plants. The reaction mechanism is classified in three different types on the basis of the affinity of the membrane complex towards its electron donor protein. The role of electrostatic interactions in forming an intermediate transient complex, as well as the structural and functional similarities of cytochrome c6 and plastocyanin are analysed from an evolutionary point of view. The proposal made is that the heme protein was first “discovered” by nature, when iron was much more abundant on the Earth's surface, and replaced by plastocyanin when copper became available because of the oxidizing conditions of the new atmosphere.