Hypoxanthine-Guanine Phosphoribosyltransferase/adenylate Kinase From Zobellia galactanivorans: A Bifunctional Catalyst for the Synthesis of Nucleoside-5′-Mono-, Di- and Triphosphates

In our search for novel biocatalysts for the synthesis of nucleic acid derivatives, we found a good candidate in a putative dual-domain hypoxanthine-guanine phosphoribosyltransferase (HGPRT)/adenylate kinase (AMPK) from Zobellia galactanivorans (ZgHGPRT/AMPK). In this respect, we report for the firs...

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Bibliographic Details
Authors: Acosta, Javier, Del Arco, Jon, Del Pozo, Maria Luisa, Herrera, Beliña, Clemente-Suárez, Vicente Javier, Berenguer, José, Hidalgo, Aurelio, Fernández-Lucas, Jesús
Format: article
Status:Versión aceptada para publicación
Publication Date:2020
Country:Colombia
Institution:Corporación Universidad de la Costa
Repository:Repositorio REDICUC
Language:English
OAI Identifier:oai:repositorio.cuc.edu.co:11323/6477
Online Access:https://hdl.handle.net/11323/6477
https://doi.org/10.3389/fbioe.2020.00677
https://repositorio.cuc.edu.co/
Access Level:Open access
Keyword:Enzymatic synthesis
Nucleotides
Phosphoribosyltransferase
Nucleoside-5cpsdummy′-monophosphate kinase
Dual domain protein
Description
Summary:In our search for novel biocatalysts for the synthesis of nucleic acid derivatives, we found a good candidate in a putative dual-domain hypoxanthine-guanine phosphoribosyltransferase (HGPRT)/adenylate kinase (AMPK) from Zobellia galactanivorans (ZgHGPRT/AMPK). In this respect, we report for the first time the recombinant expression, production, and characterization of a bifunctional HGPRT/AMPK. Biochemical characterization of the recombinant protein indicates that the enzyme is a homodimer, with high activity in the pH range 6-7 and in a temperature interval from 30 to 80°C. Thermal denaturation experiments revealed that ZgHGPRT/AMPK exhibits an apparent unfolding temperature (Tm) of 45°C and a retained activity of around 80% when incubated at 40°C for 240 min. This bifunctional enzyme shows a dependence on divalent cations, with a remarkable preference for Mg2+ and Co2+ as cofactors. More interestingly, substrate specificity studies revealed ZgHGPRT/AMPK as a bifunctional enzyme, which acts as phosphoribosyltransferase or adenylate kinase depending upon the nature of the substrate. Finally, to assess the potential of ZgHGPRT/AMPK as biocatalyst for the synthesis of nucleoside-5′-mono, di- and triphosphates, the kinetic analysis of both activities (phosphoribosyltransferase and adenylate kinase) and the effect of water-miscible solvents on enzyme activity were studied.