UGA4 gene encoding the γ-aminobutyric acid permease in Saccharomyces cerevisiae is an acid-expressed gene

Background and aims: biological processes in all organisms are controlled by environmental conditions, however, information concerning the molecular responses to external pH is scarce. In this work we studied the pH response of UGA4 gene encoding -aminolevulinic acid and -aminobutyric acid permease...

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Detalles Bibliográficos
Autores: Bermudez Moretti, Mariana, Batlle, Alcira María del C., Correa Garcia, Susana Raquel
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2001
País:Argentina
Institución:Consejo Nacional de Investigaciones Científicas y Técnicas
Repositorio:CONICET Digital (CONICET)
Idioma:inglés
OAI Identifier:oai:ri.conicet.gov.ar:11336/102043
Acceso en línea:http://hdl.handle.net/11336/102043
Access Level:acceso abierto
Palabra clave:UGA4 gene
Saccharomyces cerevisiae
Ambient pH
Transcriptional regulation
https://purl.org/becyt/ford/1.4
https://purl.org/becyt/ford/1
Descripción
Sumario:Background and aims: biological processes in all organisms are controlled by environmental conditions, however, information concerning the molecular responses to external pH is scarce. In this work we studied the pH response of UGA4 gene encoding -aminolevulinic acid and -aminobutyric acid permease in Saccharomyces cere isiae. Methods: we analyzed the effect of pH on the expression of UGA4 gene measuring -galactosidase activity in cells carrying a UGA4::lacZ fusion gene. Results: results indicate that UGA4 expression is higher at acidic pH. The expression of UGA3 and UGA35 genes, which encode two positive transcription factors, is not regulated by external pH, while the expression of UGA43 gene encoding a repressor of UGA4 transcription is dependent on pH. Using a strain lacking Uga43p we clearly showed that the effect of ambient pH on UGA4 expression is not a secondary effect of the pH regulation on UGA43. We have also demonstrated that the effect of pH can only be detected when UGA4 gene is not subject to a strong repression by Uga43p nor to GABA induction. Conclusion: here, we demonstrate that UGA4 is an acid-expressed gene. This regulation is probably mediated by Rim101p through the consensus site 5-GCCARG-3 at 237 bp preceding the UGA4 coding sequence.